The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Peruvian primrose-willow

Carolina primrose-willow

Habit Herbs often woody at base, with peeling bark. Herbs, from woody rootstock.
Stems

usually ridged, rarely succulent, profusely branched, 100–400 cm, usually villous, rarely glabrous, hairs deciduous in age, multicellular, usually tawny.

erect, subterete, 20–120 cm, branched, glabrate proximally, or strigillose, especially in distal parts, with raised strigillose lines decurrent from leaf axils mid stem.

Leaves

stipules deciduous, narrowly deltate, 1–1.5 × 0.3–0.5 mm, setaceous;

petiole 0–1.5 cm;

blade usually lanceolate, elliptic or broadly elliptic, sometimes ovate, obovate, or rounded, 2–45 × 1–10 cm, base obtuse or cuneate, rarely asymmetrical, margins entire or gland-toothed, apex usually acute or acuminate, rarely rounded and emarginate, mostly scabrid, membranous or papery, surfaces usually villous, sometimes glabrous;

bracts usually not strongly reduced.

stipules narrowly deltate, 0.3–0.5 × 0.1–0.2 mm;

petiole winged, 0.1–0.7 cm;

blade narrowly to broadly lanceolate, 4–15 × 0.3–1(–3) cm, base tapered, margins subentire to inconspicuously glandular-serrulate, apex acute to acuminate, surfaces finely strigillose, especially on abaxial veins, sometimes glabrate;

bracts narrower, reduced in size.

Inflorescences

leafy racemes, flowers solitary in distal axils;

bracteoles deciduous, usually attached near base or on lower 1/2 of ovary, sometimes on upper pedicel, subtended by reduced, glandlike stipels, ovate or lanceolate to linear, 5–20 × 1–6 mm, apex acute or short-acuminate, surfaces villous.

open, leafy racemes, flowers solitary in axils;

bracteoles lanceolate-linear or setaceous, 2–6 × 0.3–0.8 mm, attached on pedicel just proximal to base of ovary.

Flowers

sepals ovate or ovate-lanceolate, 10–23 × 4–9 mm, apex acute or short-acuminate, sometimes glandular-serrulate;

petals bright yellow, orbiculate or obovate, 10–40 × 10–40 mm, apex rarely emarginate, short-clawed;

stamens 8(or 10) in 2 unequal series, yellow, shorter filaments 1.5–4 mm, longer ones 3.5–5 mm, anthers oblong, 3–6 mm;

ovary obconic, 4- or 5-angled, sometimes subterete, 5–20 × 3–7 mm, narrowed to pedicel, usually densely villous, sometimes glabrous;

nectary disc elevated 0.3–3.2 mm on ovary apex, 4–6 mm diam., 4(or 5)-lobed, sunken, ringed by long, white hairs;

style 1.5–3.5 mm, stigma globose, 1.6–3.5 × 1.8–3.5 mm, usually as long as stamens, rarely exserted beyond them.

sepals ovate-deltate, 10–20 × 7–12 mm, abruptly acuminate or acute, inconspicuously 5–7-nerved, surfaces strigillose;

petals deep golden yellow, broadly obovate, 20–35 × 10–30 mm, apex shallowly emarginate, claw 1.5–3 mm;

stamens 8 in 2 unequal series, filaments flattened and dilated near base, epipetalous set 3.4–4.5 mm, episepalous set 4.5–5.5 mm, anthers oblong, 4–5 mm;

pollen shed in polyads;

ovary subcylindric, slightly 4-angled, 8–12(–20) mm;

nectary disc slightly elevated on ovary apex, 2–3 mm diam., 4-lobed, ringed by short hairs;

style 3–3.5 mm, stigma clavate-capitate, 2.5–3 × 2–2.5 mm, often exserted beyond anthers.

Capsules

± sharply 4- or 5-angled, 10–40 × 6–13 mm, thin-walled, irregularly dehiscent, pedicel 5–65 mm.

clavate-cylindric, subterete to obtusely 4-angled, 20–35 × 3.5–5 mm, thin walls, irregularly dehiscent, tapering to pedicel 10–40 mm.

Seeds

brown or reddish brown, oblong, 0.6–0.9 × 0.3–0.4 mm, rounded at ends, with inconspicuous raphe.

in several indistinct rows per locule, yellow-brown, oblong (appearing round), 0.5 mm, shiny, raphe 2/3 as wide as body.

2n

= 64, 80, 96, 128.

= 16.

Ludwigia peruviana

Ludwigia bonariensis

Phenology Flowering Jun–Aug(–Sep) (sometimes in any month). Flowering summer.
Habitat Wet places, ditches, drainage canals, sloughs, swales, marshy shores, wet clearings. Wet places, mainly along coastal areas, especially ditches, banks near brackish water.
Elevation 0–200[–2600] m. (0–700[–8500] ft.) 0–200[–2600] m. (0–700[–8500] ft.)
Distribution
from FNA
AL; FL; GA; NC; TX; Central America; South America; Mexico (Chiapas, Durango, Oaxaca, Puebla, Tabasco, Veracruz); West Indies [Introduced in Eurasia (India, Indonesia, Singapore, Sri Lanka), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; MS; NC; SC; TX; Mexico (Chihuahua, Durango, Puebla, Quintana Roo, Tabasco, Tamaulipas, Veracruz); South America (Argentina, Bolivia, Brazil, Paraguay, Uruguay)
[WildflowerSearch map]
[BONAP county map]
Discussion

Ludwigia peruviana is sometimes cultivated and naturalized, which may account for occurrences in North America and Eurasia. Except in the Amazon basin, where it is known only from few collections in western Amazonia, and in northeastern Brazil, where it is scarce, L. peruviana is common throughout its range, and may behave as a weed, especially along slow-flowing canals and drainage ditches. Ludwigia peruviana is also naturalized at scattered localities in Asia (P. H. Raven 1963[1964]) and around Sydney, Australia. The earliest collection from Asia is labeled “ex horto bot. Bogoriensi Javae misit 1869” (Raven). It also occurs locally in the Nilgiri Mountains of southwest India and in Sri Lanka, as well as in scattered locations in Bangka, Java, Malaysia, and Sumatra from sea level to 1000 m (Raven 1963[1964], 1978).

Jussiaea grandiflora Ruíz & Pavon, a synonym for J. peruviana Linnaeus, appeared in 1830, not in 1802 (P. A. Munz 1942; P. H. Raven 1963[1964]); it is a later homonym of J. grandiflora Michaux (1803), as reported in W. Greuter and T. Raus (1987). Jussiaea hirta (Linnaeus) Swartz is an illegitimate homonym and J. hirta (Linnaeus) Vahl is an illegitimate isonym; both pertain here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Myrtocarpus Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Macrocarpon
Sibling taxa
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
L. alata, L. alternifolia, L. arcuata, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
Synonyms Jussiaea peruviana, J. hirsuta, J. macrocarpa, J. mollis, J. peruviana var. glaberrima, J. speciosa, J. sprengeri, L. hirta, Oenothera hirta Jussiaea bonariensis, J. neglecta, J. suffruticosa var. bonariensis
Name authority (Linnaeus) H. Hara: J. Jap. Bot. 28: 293. (1953) (Micheli) H. Hara: J. Jap. Bot. 28: 291. (1953)
Web links