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Photo is of parent taxon

floating primrose-willow, floating water primrose, Matted water primrose

creeping water primrose, floating primrose-willow, marsh purslane

Habit Herbs or emergent aquatics, rooting at nodes, sometimes with fleshy, white pneumatophores at submerged nodes.
Stems

glabrous or sparsely villous.

floating or creeping and ascending to erect, terete, 10–100(–300) cm, simple or branched, glabrous or sparsely to densely villous, hairs sometimes viscid on emergent distal stem.

Leaves

alternate or fascicled;

stipules symmetrical;

petioles of basal leaves 0.3–0.8 cm, those of distal leaves 0.2–2.5 cm;

blade 0.8–4(–8.5) cm, apex sometimes glandular-mucronate, surfaces shiny, glabrous, rarely ciliate or minutely pellucid-punctate.

stipules broadly ovate-deltate, 0.6–1.6 × 0.4–1 mm, succulent, apex acute or obtuse, gland-tipped, rarely divided into 3 parts;

petiole flattened or narrowly winged, 0.2–6 cm;

blade narrowly oblong or elliptic to ovate, broadly obovate, or orbiculate, (0.4–)1–10 × 0.4–4 cm, base narrowly cuneate or attenuate, margins entire, apex obtuse or rounded to acute, sometimes mucronate or glandular-mucronate, surfaces of floating leaves glabrous, those of emergent leaves glabrous to sparsely or densely strigillose at least adaxially;

bracts scarcely reduced.

Inflorescences

on emergent stems sometimes in leafy racemes, flowers solitary in leaf axils;

bracteoles (rarely absent), deltate, squamate, 0.5–1 × 0.5–1 mm, apex acute, attached near base or on lower 1/2 of ovary.

Flowers

anthers on short filaments 0.5–1.1 mm, those on long filaments 0.8–1.2 mm;

ovary 6–14 mm, apex truncate, glabrous or with scattered hairs, stigma usually as long as anthers, rarely exserted beyond them.

sepals narrowly deltate or lanceolate, 3–12 × 1.5–4 mm, apex acute or acuminate, surfaces glabrous or sparsely to densely hirtellous;

petals yellow, obpyramidal,7–24 × 4–13 mm, apex mucronate or emarginate, up-curved;

stamens 10(or 12), in 2 unequal series, bright yellow, filaments suberect or reflexed, shorter ones 1.4–4.2 mm, longer ones (1.9–)3.3–6 mm, anthers oblong, 0.5–2.2 mm;

ovary subcylindric or truncate, 6–20 × 1.5–3 mm, apex somewhat broader, glabrous or sparsely to densely hirtellous;

nectary disc slightly raised on ovary apex, 2–2.5 mm diam., lobed, glabrous or fringed with long hairs;

style (1.9–)2.4–7.3 mm, glabrous or sparsely to densely hirtellous on proximal 1/2, stigma flattened-globose, 0.9–1.2 × 1–2.5 mm, sometimes shallowly or deeply 5-lobed, as long as or exserted beyond anthers.

Capsules

10–17(–25) × 2–3 mm, pedicel 10–35 mm.

cylindric, subterete to obscurely 5-angled, straight or curved, 10–40 × 2–4 mm, with thick woody walls, irregularly and tardily dehiscent, pedicel 7–60(–90) mm.

Seeds

7–14 per locule.

embedded in elongated piece of endocarp, 1–1.5 × 0.9–1.3 mm.

2n

= 16.

= 16.

Ludwigia peploides subsp. peploides

Ludwigia peploides

Phenology Flowering summer–early fall.
Habitat Wet places, along slow-moving rivers, streams, canals, ditches, often growing into main channels as aquatic weeds.
Elevation 0–900[–3000] m. (0–3000[–9800] ft.)
Distribution
from FNA
AZ; CA; NM; OR; TX; Mexico; Central America; South America; West Indies (Cuba); Pacific Islands (Galapagos Islands) [Introduced elsewhere in Pacific Islands (Society Islands), Australia]
[BONAP county map]
from USDA
United States; Mexico; Central America; South America; West Indies (Cuba); Asia (China); Pacific Islands (Galapagos Islands) [Introduced in Europe (France), elsewhere in the Pacific Islands (New Zealand, Society Islands), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (3 in the flora).

Ludwigia peploides consists of four subspecies more or less well defined geographically and morphologically, with three present in the flora area: subsp. glabrescens, subsp. montevidensis, and subsp. peploides (P. H. Raven 1963[1964]); these subspecies have ranges that are mostly distinct. Subspecies peploides has a wide distribution in the New World, from the southern United States south to Argentina. Subspecies glabrescens is widespread in eastern United States. Subspecies montevidensis occurs primarily in southern South America and scattered (probably introduced) in the southern United States, Australia, France, and New Zealand. Subspecies peploides and montevidensis occur together locally in California and Louisiana, where subsp. montevidensis is introduced. The ranges of subsp. glabrescens and peploides come together in Texas.

Subspecies stipulacea (Ohwi) P. H. Raven is known from eastern Asia (e China).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually densely villous, rarely sparsely so; leaf blades not shiny, apices glandular-mucronate; capsules (20–)24–32 mm.
subsp. montevidensis
1. Stems glabrous or sparely villous; leaf blades shiny, apices usually eglandular-mucronate; capsules 10–40 mm.
→ 2
2. Principal leaf blades 0.8–4(–8.5) cm; petioles 0.2–2.5 cm; pedicels 10–35 mm; capsules 10–17(–25) mm; seeds 7–14 per locule.
subsp. peploides
2. Principal leaf blades (2–)4–10 cm; petioles 0.7–6 cm; pedicels 35–90 mm; capsule 25–40 mm; seeds 16–18 per locule.
subsp. glabrescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Jussiaea > Ludwigia peploides Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Jussiaea
Sibling taxa
L. peploides subsp. glabrescens, L. peploides subsp. montevidensis
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
Subordinate taxa
L. peploides subsp. glabrescens, L. peploides subsp. montevidensis, L. peploides subsp. peploides
Synonyms Jussiaea californica, J. diffusa var. californica, J. repens var. californica Jussiaea peploides, J. repens var. peploides, L. adscendens var. peploides
Name authority unknown (Kunth) P. H. Raven: Reinwardtia 6: 393. (1964)
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