Ludwigia peploides subsp. montevidensis |
Ludwigia peploides |
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floating primrose-willow, floating water primrose |
creeping water primrose, floating primrose-willow, marsh purslane |
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Habit | Herbs or emergent aquatics, rooting at nodes, sometimes with fleshy, white pneumatophores at submerged nodes. | |||||||||
Stems | usually densely villous, rarely sparsely so, hairs often viscid when fresh, or glabrate on submerged stems. |
floating or creeping and ascending to erect, terete, 10–100(–300) cm, simple or branched, glabrous or sparsely to densely villous, hairs sometimes viscid on emergent distal stem. |
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Leaves | alternate, sometimes fascicled; stipules often asymmetrical; petioles of basal leaves (0.5–)0.8–1.6 cm, those of distal leaves 0.5–2.8 cm; blade (0.4–)1–6(–9.5) cm, apex glandular-mucronate, surfaces not shiny, usually densely hirtellous, rarely glabrous abaxially. |
stipules broadly ovate-deltate, 0.6–1.6 × 0.4–1 mm, succulent, apex acute or obtuse, gland-tipped, rarely divided into 3 parts; petiole flattened or narrowly winged, 0.2–6 cm; blade narrowly oblong or elliptic to ovate, broadly obovate, or orbiculate, (0.4–)1–10 × 0.4–4 cm, base narrowly cuneate or attenuate, margins entire, apex obtuse or rounded to acute, sometimes mucronate or glandular-mucronate, surfaces of floating leaves glabrous, those of emergent leaves glabrous to sparsely or densely strigillose at least adaxially; bracts scarcely reduced. |
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Inflorescences | on emergent stems sometimes in leafy racemes, flowers solitary in leaf axils; bracteoles (rarely absent), deltate, squamate, 0.5–1 × 0.5–1 mm, apex acute, attached near base or on lower 1/2 of ovary. |
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Flowers | anthers on short filaments (0.7–)0.9–1.8 mm, those on long filaments (0.8–)1.1–2.2 mm; ovary 6–10 mm, apex truncate, densely hirtellous, sometimes only on apical 1/2, stigma usually as long as anthers, rarely exserted beyond them. |
sepals narrowly deltate or lanceolate, 3–12 × 1.5–4 mm, apex acute or acuminate, surfaces glabrous or sparsely to densely hirtellous; petals yellow, obpyramidal,7–24 × 4–13 mm, apex mucronate or emarginate, up-curved; stamens 10(or 12), in 2 unequal series, bright yellow, filaments suberect or reflexed, shorter ones 1.4–4.2 mm, longer ones (1.9–)3.3–6 mm, anthers oblong, 0.5–2.2 mm; ovary subcylindric or truncate, 6–20 × 1.5–3 mm, apex somewhat broader, glabrous or sparsely to densely hirtellous; nectary disc slightly raised on ovary apex, 2–2.5 mm diam., lobed, glabrous or fringed with long hairs; style (1.9–)2.4–7.3 mm, glabrous or sparsely to densely hirtellous on proximal 1/2, stigma flattened-globose, 0.9–1.2 × 1–2.5 mm, sometimes shallowly or deeply 5-lobed, as long as or exserted beyond anthers. |
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Capsules | (20–)24–32 × 2–4 mm, pedicel 7–38(–60) mm. |
cylindric, subterete to obscurely 5-angled, straight or curved, 10–40 × 2–4 mm, with thick woody walls, irregularly and tardily dehiscent, pedicel 7–60(–90) mm. |
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Seeds | 10–15 per locule. |
embedded in elongated piece of endocarp, 1–1.5 × 0.9–1.3 mm. |
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2n | = 16 (32). |
= 16. |
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Ludwigia peploides subsp. montevidensis |
Ludwigia peploides |
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Phenology | Flowering summer-early fall. | |||||||||
Habitat | Wet places, along slow-moving rivers, streams, canals, ditches, often growing into main channels as aquatic weeds. | |||||||||
Elevation | 0–500[–2000] m. (0–1600[–6600] ft.) | |||||||||
Distribution |
CA; LA; South America (Argentina, Brazil, Chile, Peru, Uruguay) [Introduced in North America; introduced also in Europe (France), Pacific Islands (New Zealand), Australia] |
United States; Mexico; Central America; South America; West Indies (Cuba); Asia (China); Pacific Islands (Galapagos Islands) [Introduced in Europe (France), elsewhere in the Pacific Islands (New Zealand, Society Islands), Australia]
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Discussion | In the flora area, subsp. montevidensis is introduced in California (P. H. Raven 1963c), where it was first collected in 1906 (El Dorado County, Rixford s.n., CAS), and in Louisiana. Subspecies montevidensis occasionally forms masses of vegetation that can obstruct water flow and navigation in California and elsewhere. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (3 in the flora). Ludwigia peploides consists of four subspecies more or less well defined geographically and morphologically, with three present in the flora area: subsp. glabrescens, subsp. montevidensis, and subsp. peploides (P. H. Raven 1963[1964]); these subspecies have ranges that are mostly distinct. Subspecies peploides has a wide distribution in the New World, from the southern United States south to Argentina. Subspecies glabrescens is widespread in eastern United States. Subspecies montevidensis occurs primarily in southern South America and scattered (probably introduced) in the southern United States, Australia, France, and New Zealand. Subspecies peploides and montevidensis occur together locally in California and Louisiana, where subsp. montevidensis is introduced. The ranges of subsp. glabrescens and peploides come together in Texas. Subspecies stipulacea (Ohwi) P. H. Raven is known from eastern Asia (e China). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||
Parent taxa | ||||||||||
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Synonyms | Jussiaea montevidensis, J. repens var. montevidensis, L. adscendens var. montevidensis, L. peploides var. montevidensis | Jussiaea peploides, J. repens var. peploides, L. adscendens var. peploides | ||||||||
Name authority | (Sprengel) P. H. Raven: Reinwardtia 6: 395. (1964) | (Kunth) P. H. Raven: Reinwardtia 6: 393. (1964) | ||||||||
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