The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

creeping water primrose, floating primrose-willow, marsh purslane

shrubby primrose-willow, shrubby primrose-willow or seedbox

Habit Herbs or emergent aquatics, rooting at nodes, sometimes with fleshy, white pneumatophores at submerged nodes. Herbs with 1–3 rhizomes 0.6–5.5 × 0.2–0.5 mm, often branched, glabrous or densely hirtellous, sometimes also forming stolons 8–80 cm, 1.1–2.2 mm thick, branched, glabrous or sparsely to densely hirtellous.
Stems

floating or creeping and ascending to erect, terete, 10–100(–300) cm, simple or branched, glabrous or sparsely to densely villous, hairs sometimes viscid on emergent distal stem.

erect, slightly ridged, unbranched or slightly branched, (16–)30–90 cm, glabrous or strigillose to hirtellous, especially on distal parts.

Leaves

stipules broadly ovate-deltate, 0.6–1.6 × 0.4–1 mm, succulent, apex acute or obtuse, gland-tipped, rarely divided into 3 parts;

petiole flattened or narrowly winged, 0.2–6 cm;

blade narrowly oblong or elliptic to ovate, broadly obovate, or orbiculate, (0.4–)1–10 × 0.4–4 cm, base narrowly cuneate or attenuate, margins entire, apex obtuse or rounded to acute, sometimes mucronate or glandular-mucronate, surfaces of floating leaves glabrous, those of emergent leaves glabrous to sparsely or densely strigillose at least adaxially;

bracts scarcely reduced.

alternate;

stipules deltate, 0.25–0.45 × 0.15–0.4 mm;

rhizomes: sessile, blades minute, appressed, and scalelike, oblate or suborbiculate, 0.3–0.6 × 0.6–0.9 mm;

stolons: petiole 0.1–0.6 cm, blade oblong or oblanceolate-elliptic to spatulate, 0.4–3.5 × 0.2–1.5 cm;

main stem: sessile, blade lanceolate-elliptic or lanceolate-linear to linear, 2.5–9.5 × (0.1–)0.3–0.9 cm, proximal ones shorter and often oblong or oblong-lanceolate, base rounded or obtuse, margins entire with obscure hydathodal glands, apex acuminate to acute, surfaces glabrous or, sometimes, pilose on proximal blades;

bracts very reduced.

Inflorescences

on emergent stems sometimes in leafy racemes, flowers solitary in leaf axils;

bracteoles (rarely absent), deltate, squamate, 0.5–1 × 0.5–1 mm, apex acute, attached near base or on lower 1/2 of ovary.

densely clustered, terminal racemes or spikes, 1–5(–12) cm;

bracteoles attached at base of ovary or on pedicel distally, narrowly lanceolate, 3.5–5(–6) × (1.2–)1.4–2 mm, surfaces usually glabrous, sometimes pilose abaxially.

Flowers

sepals narrowly deltate or lanceolate, 3–12 × 1.5–4 mm, apex acute or acuminate, surfaces glabrous or sparsely to densely hirtellous;

petals yellow, obpyramidal,7–24 × 4–13 mm, apex mucronate or emarginate, up-curved;

stamens 10(or 12), in 2 unequal series, bright yellow, filaments suberect or reflexed, shorter ones 1.4–4.2 mm, longer ones (1.9–)3.3–6 mm, anthers oblong, 0.5–2.2 mm;

ovary subcylindric or truncate, 6–20 × 1.5–3 mm, apex somewhat broader, glabrous or sparsely to densely hirtellous;

nectary disc slightly raised on ovary apex, 2–2.5 mm diam., lobed, glabrous or fringed with long hairs;

style (1.9–)2.4–7.3 mm, glabrous or sparsely to densely hirtellous on proximal 1/2, stigma flattened-globose, 0.9–1.2 × 1–2.5 mm, sometimes shallowly or deeply 5-lobed, as long as or exserted beyond anthers.

sepals ascending, pale green or white adaxially, broadly ovate-deltate, 2.3–3.5(–4) × 2.3–3.2(–3.8) mm, margins entire, apex acuminate, surfaces glabrous;

petals 0;

filaments yellow or cream, 1.2–2 mm, distinctly dilated toward base, anthers 0.7–1(–1.3) × 0.5–0.7 mm;

pollen shed in tetrads;

ovary broadly obovoid or cup-shaped, 2.2–3 × 2.3–3.3 mm;

nectary disc elevated 0.5–0.6 mm on ovary apex, pale yellow, 1.8–3.1 mm diam., obscurely 4-lobed, glabrous;

style pale yellow, 0.9–1.7 mm, glabrous, stigma pale green to white, globose to capitate, 0.4–0.8 × 0.5–0.8 mm, distinctly 4-lobed, not exserted beyond anthers.

Capsules

cylindric, subterete to obscurely 5-angled, straight or curved, 10–40 × 2–4 mm, with thick woody walls, irregularly and tardily dehiscent, pedicel 7–60(–90) mm.

broadly obpyramidal, angles rounded, sometimes subspherical, 2.5–4.3 ×2.5–4.5(–5) mm, hard-walled, dehiscent by apical ring, pedicel 0.5–1.5(–2) mm.

Seeds

embedded in elongated piece of endocarp, 1–1.5 × 0.9–1.3 mm.

brown, elliptic-oblong, curved on both ends, 0.5–0.6 × 0.2–0.3 mm, surface cells ± isodiametric.

2n

= 16.

= 32.

Ludwigia peploides

Ludwigia suffruticosa

Phenology Flowering May–Sep.
Habitat Sandy ditches, marshes, wet meadows, limestone sinks, cypress swamps, moist pinelands.
Elevation 0–150 m. (0–500 ft.)
Distribution
from USDA
United States; Mexico; Central America; South America; West Indies (Cuba); Asia (China); Pacific Islands (Galapagos Islands) [Introduced in Europe (France), elsewhere in the Pacific Islands (New Zealand, Society Islands), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; NC; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (3 in the flora).

Ludwigia peploides consists of four subspecies more or less well defined geographically and morphologically, with three present in the flora area: subsp. glabrescens, subsp. montevidensis, and subsp. peploides (P. H. Raven 1963[1964]); these subspecies have ranges that are mostly distinct. Subspecies peploides has a wide distribution in the New World, from the southern United States south to Argentina. Subspecies glabrescens is widespread in eastern United States. Subspecies montevidensis occurs primarily in southern South America and scattered (probably introduced) in the southern United States, Australia, France, and New Zealand. Subspecies peploides and montevidensis occur together locally in California and Louisiana, where subsp. montevidensis is introduced. The ranges of subsp. glabrescens and peploides come together in Texas.

Subspecies stipulacea (Ohwi) P. H. Raven is known from eastern Asia (e China).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ludwigia suffruticosa is distinctive by virtue of its highly condensed inflorescence and sessile leaves. It also differs from other species in sect. Isnardia in that it perennates mainly by underground rhizomes. This apetalous species has showy bracts and attracts many insects, including bumblebees, wasps, and honeybees (C. I. Peng 1989). Its center of distribution is in Florida, extending along the coastal plain barely to Alabama on the west and barely to southern North Carolina on the northeast. Recent reports of this species from Mississippi and from Mexico (Chiapas and Oaxaca) have not been confirmed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually densely villous, rarely sparsely so; leaf blades not shiny, apices glandular-mucronate; capsules (20–)24–32 mm.
subsp. montevidensis
1. Stems glabrous or sparely villous; leaf blades shiny, apices usually eglandular-mucronate; capsules 10–40 mm.
→ 2
2. Principal leaf blades 0.8–4(–8.5) cm; petioles 0.2–2.5 cm; pedicels 10–35 mm; capsules 10–17(–25) mm; seeds 7–14 per locule.
subsp. peploides
2. Principal leaf blades (2–)4–10 cm; petioles 0.7–6 cm; pedicels 35–90 mm; capsule 25–40 mm; seeds 16–18 per locule.
subsp. glabrescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Jussiaea Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Isnardia
Sibling taxa
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. virgata
Subordinate taxa
L. peploides subsp. glabrescens, L. peploides subsp. montevidensis, L. peploides subsp. peploides
Synonyms Jussiaea peploides, J. repens var. peploides, L. adscendens var. peploides Isnardia suffruticosa
Name authority (Kunth) P. H. Raven: Reinwardtia 6: 393. (1964) Walter: Fl. Carol., 90. (1788)
Web links