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Mexican primrose-willow

creeping water primrose, floating primrose-willow, marsh purslane

Habit Herbs (robust)or shrubs, herbs tap-rooted, often woody at base, with peeling bark. Herbs or emergent aquatics, rooting at nodes, sometimes with fleshy, white pneumatophores at submerged nodes.
Stems

erect to spreading, terete or sometimes ridged, 60–250(–400) cm, densely branched, densely villous to glabrate, especially near base.

floating or creeping and ascending to erect, terete, 10–100(–300) cm, simple or branched, glabrous or sparsely to densely villous, hairs sometimes viscid on emergent distal stem.

Leaves

stipules deltate, 0.5–0.8 × 0.6–0.7 mm, fleshy;

petiole 0–1 cm;

blade linear to oblong or oblanceolate, sometimes narrowly ovate, 0.7–14.5 × 0.1–4 cm, base tapered, margins entire, apex acute to acuminate, surfaces ± densely villous or strigillose;

bracts not or scarcely reduced.

stipules broadly ovate-deltate, 0.6–1.6 × 0.4–1 mm, succulent, apex acute or obtuse, gland-tipped, rarely divided into 3 parts;

petiole flattened or narrowly winged, 0.2–6 cm;

blade narrowly oblong or elliptic to ovate, broadly obovate, or orbiculate, (0.4–)1–10 × 0.4–4 cm, base narrowly cuneate or attenuate, margins entire, apex obtuse or rounded to acute, sometimes mucronate or glandular-mucronate, surfaces of floating leaves glabrous, those of emergent leaves glabrous to sparsely or densely strigillose at least adaxially;

bracts scarcely reduced.

Inflorescences

open, leafy racemes, flowers solitary in axils, presentation often radial;

bracteoles ovate, 3–8 × 1.4–4 mm, apex acuminate, attached near base of ovary.

on emergent stems sometimes in leafy racemes, flowers solitary in leaf axils;

bracteoles (rarely absent), deltate, squamate, 0.5–1 × 0.5–1 mm, apex acute, attached near base or on lower 1/2 of ovary.

Flowers

sepals lanceolate to ovate, (6–)8–13 × 3–7 mm, apex acuminate, surfaces strigillose adaxially;

petals bright yellow, fan-shaped, (5–)10–20 × 5–20 mm, apex sometimes shallowly notched;

stamens 8 in 2 subequal series, yellowish white, filaments spreading, 4–6 mm, anthers oblong, 2.5–5 × 1–2 mm;

pollen shed in tetrads or sometimes polyads;

ovary cylindric, 4-angled, sometimes slightly twisted, 8–22 × 1–3 mm;

nectary disc elevated 0.4–0.5 mm on ovary apex, 1–2.4 mm diam., with 4 white-pubescent sunken lobes opposite petals;

style 2.5–3.5 mm, stigma capitate-globose, 1.8–3.5 × 2.5–3.5 mm, surrounded by anthers and pollen shed directly on it.

sepals narrowly deltate or lanceolate, 3–12 × 1.5–4 mm, apex acute or acuminate, surfaces glabrous or sparsely to densely hirtellous;

petals yellow, obpyramidal,7–24 × 4–13 mm, apex mucronate or emarginate, up-curved;

stamens 10(or 12), in 2 unequal series, bright yellow, filaments suberect or reflexed, shorter ones 1.4–4.2 mm, longer ones (1.9–)3.3–6 mm, anthers oblong, 0.5–2.2 mm;

ovary subcylindric or truncate, 6–20 × 1.5–3 mm, apex somewhat broader, glabrous or sparsely to densely hirtellous;

nectary disc slightly raised on ovary apex, 2–2.5 mm diam., lobed, glabrous or fringed with long hairs;

style (1.9–)2.4–7.3 mm, glabrous or sparsely to densely hirtellous on proximal 1/2, stigma flattened-globose, 0.9–1.2 × 1–2.5 mm, sometimes shallowly or deeply 5-lobed, as long as or exserted beyond anthers.

Capsules

cylindric to clavate-cylindric, subterete to ± 4-angled, 17–50 × 2.5–8 mm, thin-walled, irregularly dehiscent by 4–8 linear valves splitting from apex, short-villous, pedicel 5–25 mm.

cylindric, subterete to obscurely 5-angled, straight or curved, 10–40 × 2–4 mm, with thick woody walls, irregularly and tardily dehiscent, pedicel 7–60(–90) mm.

Seeds

in several indistinct rows per locule, broad-cylindric with rounded ends, 0.6–0.9 × 0.5–0.8 mm, raphe inflated and nearly equal to seed body.

embedded in elongated piece of endocarp, 1–1.5 × 0.9–1.3 mm.

2n

= [16], 32, 48.

= 16.

Ludwigia octovalvis

Ludwigia peploides

Phenology Flowering summer–early fall.
Habitat Wet or moist places, along coasts, streams, ditches, swamps, often near disturbance or cultivation.
Elevation 0–300[–2200] m. (0–1000[–7200] ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; Mexico; Central America; South America; West Indies; s Asia; e Asia (Burma, China, India, Japan, Thailand, Vietnam); Africa; Indian Ocean Islands (Comoros Islands, Madagascar); Pacific Islands (New Caledonia, Papua New Guinea, Philippines)
[WildflowerSearch map]
[BONAP county map]
from USDA
United States; Mexico; Central America; South America; West Indies (Cuba); Asia (China); Pacific Islands (Galapagos Islands) [Introduced in Europe (France), elsewhere in the Pacific Islands (New Zealand, Society Islands), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Ludwigia octovalvis is perhaps the most widespread species of Ludwigia worldwide and exhibits a very complex pattern of morphological and ecological variation, correlated only in part with multiple ploidy levels; this complexity is reflected in its extensive synonymy. P. H. Raven (1963[1964]), P. A. Munz (1942, 1965), and others have proposed formal classifications to account for this variation, with mixed results and additional study using more powerful analytical tools is clearly needed in order to develop a more stable classification. In the absence of better understanding, and despite some correlated patterns of morphological and geographical variation on a global scale, this treatment does not recognize infraspecific taxa.

Jussiaea hirsuta Velloso, J. suffruticosa var. angustifolia Chodat & Hassler, J. velutina Kunze, and Ludwigia suffruticosa (Linnaeus) M. Gómez are later homonyms; these four names pertain here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (3 in the flora).

Ludwigia peploides consists of four subspecies more or less well defined geographically and morphologically, with three present in the flora area: subsp. glabrescens, subsp. montevidensis, and subsp. peploides (P. H. Raven 1963[1964]); these subspecies have ranges that are mostly distinct. Subspecies peploides has a wide distribution in the New World, from the southern United States south to Argentina. Subspecies glabrescens is widespread in eastern United States. Subspecies montevidensis occurs primarily in southern South America and scattered (probably introduced) in the southern United States, Australia, France, and New Zealand. Subspecies peploides and montevidensis occur together locally in California and Louisiana, where subsp. montevidensis is introduced. The ranges of subsp. glabrescens and peploides come together in Texas.

Subspecies stipulacea (Ohwi) P. H. Raven is known from eastern Asia (e China).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually densely villous, rarely sparsely so; leaf blades not shiny, apices glandular-mucronate; capsules (20–)24–32 mm.
subsp. montevidensis
1. Stems glabrous or sparely villous; leaf blades shiny, apices usually eglandular-mucronate; capsules 10–40 mm.
→ 2
2. Principal leaf blades 0.8–4(–8.5) cm; petioles 0.2–2.5 cm; pedicels 10–35 mm; capsules 10–17(–25) mm; seeds 7–14 per locule.
subsp. peploides
2. Principal leaf blades (2–)4–10 cm; petioles 0.7–6 cm; pedicels 35–90 mm; capsule 25–40 mm; seeds 16–18 per locule.
subsp. glabrescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Macrocarpon Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Jussiaea
Sibling taxa
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
Subordinate taxa
L. peploides subsp. glabrescens, L. peploides subsp. montevidensis, L. peploides subsp. peploides
Synonyms Oenothera octovalvis, Jussiaea calycina, J. clavata, J. frutescens, J. haenkeana, J. hirta, J. ligustrifolia, J. occidentalis, J. octofila, J. octonervia, J. octonervia var. sessiliflora, J. octovalvis, J. parviflora, J. peruviana var. octofila, J. pubescens, J. sagrana, J. salicifolia, J. scabra, J. suffruticosa, J. suffruticosa var. ligustrifolia, J. suffruticosa var. linearifolia, J. suffruticosa var. octofila, J. suffruticosa subsp. octonervia, J. suffruticosa var. octonervia, J. suffruticosa var. sessiliflora, J. suffruticosa var. sintenisii, J. venosa, J. villosa, L. octovalvis var. ligustrifolia, L. octovalvis var. octofila, L. octovalvis subsp. sessiliflora, L. octovalvis var. sessiliflora, L. pubescens, L. pubescens var. ligustrifolia, L. pubescens var. linearifolia, L. pubescens var. sessiliflora, L. sagrana Jussiaea peploides, J. repens var. peploides, L. adscendens var. peploides
Name authority (Jacquin) P. H. Raven: Kew Bull. 15: 476. (1962) (Kunth) P. H. Raven: Reinwardtia 6: 393. (1964)
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