Ludwigia lanceolata |
Ludwigia hirtella |
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lanceleaf primrose-willow |
hairy seedbox, Rafinesque's seedbox, spindleroot |
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Habit | Herbs slender, with well-developed aerenchyma when base submerged, often forming stolons 10–40 cm, 2–3 mm thick, stolons with widely spaced leaves. | |
Roots | fusiform, somewhat thickened, often fascicled. |
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Stems | erect, subterete or slightly ridged, well branched distally, 45–100 cm, glabrous, with raisedlines decurrent from leaf axils. |
subterete to angled, with raised lines or narrow wings decurrent from leaf axils, (30–)50–120 cm, well branched in distal 1/2, densely erect-hirsute to sometimes glabrous. |
Leaves | alternate; stipules ovate to very broadly ovate, 0.2–0.5 ×0.1–0.3 mm, succulent; stolons: petiole winged, 0.2–1 cm, blade orbiculate or elliptic to broadly elliptic, 0.5–2.7 × 0.6–1.2 cm, base attenuate, apex rounded to acute; main stem: petiole winged, 0.1–0.5 cm, blade elliptic, oblanceolate, or narrowly oblanceolate to sublinear, 2–7.5 × 0.2–0.8(–1.4) cm, base narrowly cuneate, sometimes attenuate, margins entire with minute hydathodal glands, apex acute to narrowly acute, leaves on side branches much reduced; bracts sublinear, moderately reduced. |
stipules narrowly deltate, 0.05–0.15 × 0.05–0.1 mm; sessile; blade closely appressed to stem, lanceolate to ovate-oblong, 1.4–5.5 × 0.4–1.2 cm, base attenuate, margins entire, apex acute to rounded, surfaces glabrous or densely erect-hirsute; bracts usually very reduced, sublinear. |
Inflorescences | open spikes, flowers solitary in distal leaf axils; bracteoles attached in opposite pairs at or slightly distal to base of ovary, ovate-elliptic to very narrowly elliptic, 1.5–4.3 × 0.4–1.4 mm, often with a swollen base, margins minutely papillose, apex subacute. |
leafy racemes, flowers solitary in leaf axils; bracteoles attached in subopposite pairs just proximal to ovary base, lanceolate-linear, 2.5–7 × 0.5–1.5 mm, margins entire, apex acute, surfaces hirtellous. |
Flowers | sepals ascending, pale green, broadly ovate-deltate, 1.5–2.5 × 1.8–3.3 mm, margins minutely papillose, apex usually acute, rarely acuminate, surfaces glabrous; petals 0; filaments nearly translucent, 1–1.4 mm, base dilated, anthers 0.4–0.6(–0.8) × 0.5–0.6 mm; pollen shed in tetrads; ovary broadly obovoid or cup-shaped, 2.5–3.5 × 2.2–3.2 mm; nectary disc elevated 0.4–0.6 mm on ovary apex, yellowish green, 1.8–2.6 mm diam., 4-lobed, glabrous; style yellowish green, 0.5–0.7 mm, glabrous, stigma yellowish green, broadly capitate to subglobose, 0.3–0.5 × 0.7–1 mm, not exserted beyond anthers. |
sepals narrowly ovate-deltate, (5.4–)6–12 × 2.2–4.5(–5) mm, margins entire, apex acute, surfaces hirtellous or, sometimes, glabrous; petals cordate, 12–14 × 13–14 mm, base attenuate, apex emarginate; filaments opaque white, awl-shaped, 2–5.5 mm, anthers 1.4–2.6 × 0.4–0.8 mm; ovary subcuboid or globose, 3–4 × 3–4 mm; nectary disc elevated, domed, 1–1.4 mm diam., prominently 4-lobed, ringed with stiff, spreading hairs or glabrous; style 1.3–4.5(–5.5) mm, glabrous, stigma capitate to hemispherical, 0.9–1.5 × 0.9–2(–2.5) mm, often shallowly 4-lobed, as long as stamens, sometimes exserted beyond them. |
Capsules | obpyramidal, sharply 4-angled and 4-winged, wings 0.3–0.7mm wide, 3.5–5 × 2.5–4.5 mm, hard-walled, dehiscent by apical ring, pedicel 0.1–0.5 mm. |
subcuboid to squarish globose, 4–6 × 4–5 mm, 4-angled, often also 4-winged, wings 0.3–1.1 mm wide, pedicel 3–10 mm. |
Seeds | light brown, narrowly oblong with constricted ends, 0.6–0.8 ×0.2–0.3 mm, surface cells nearly isodiametric. |
light brown, oblong to reniform, 0.6–0.7 × 0.3–0.4 mm, surface cells elongate transversely to seed length. |
2n | = 32. |
= 16. |
Ludwigia lanceolata |
Ludwigia hirtella |
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Phenology | Flowering Jun–Oct. | Flowering summer–early fall. |
Habitat | Ditches, low meadows, cypress swamps, moist pinelands, edges of pocosins, sandy peaty soil. | Boggy depressions, roadside ditches, margins of boggy streams. |
Elevation | 0–100 m. (0–300 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
FL; GA; NC; SC
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AL; AR; DC; DE; FL; GA; KY; LA; MD; MS; NC; NJ; OK; SC; TN; TX; VA
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Discussion | Ludwigia lanceolata is fairly uncommon, with scattered populations occurring along the Atlantic coast of southern North Carolina, South Carolina, eastern and southern Georgia, and peninsular Florida. It reaches its western limit in the central panhandle of Florida. C. I. Peng (1988, 1989) found this tetraploid species to be interfertile with other tetraploid species in the section, but few natural hybrids are found, perhaps due to persistent autogamy and habitat specialization. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A name often found in synonymy with Ludwigia hirtella is a later homonym: L. hirsuta Pursh 1813, not Desrousseaux 1789. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Isnardia lanceolata | Isnardia hirsuta var. permollis, I. hirtella, L. permollis |
Name authority | Elliott: Sketch Bot. S. Carolina 1: 213. (1817) | Rafinesque: Med. Repos., hexade 2, 5: 358. (1808) |
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