Ludwigia glandulosa subsp. brachycarpac.
Onagraceae subfam. ludwigioideae
rarely reddish green, 10–55(–90) cm.
petiole 0–1 cm, blades linear-elliptic to linear, sometimes very narrowly elliptic, those on main axis 3–5(–7) × 0.3–0.5(–1) cm, those on branches 0.8–3.6 × 0.2–0.3(–0.8) cm.
stipules present.
brac-teoles attached at base of ovary, 0.4–0.8 × 0.1–0.2 mm.
sepals 1.1–1.9 × 1–1.8 mm, apex acute or short-acuminate;
nectary disc obscurely, minutely papillose;
style 0.4–0.8 mm, stigma 0.2–0.3 mm diam.
floral tube absent;
sepals (3 or)4 or 5(–7), persistent at ovary apex after dehiscence of other floral parts;
petals yellow or white, rarely absent.
obscurely 4-angled, 2–5 × 1.3–2 mm, pedicel 0–0.2 mm.
0.6–0.8 × 0.3–0.4 mm, surface cells elongate transversely to seed length.
Ludwigia glandulosa subsp. brachycarpac.
Onagraceae subfam. ludwigioideae
Subspecies brachycarpa grows along the Gulf Coast from southwestern Louisiana to Nueces County, Texas, and more sporadically northward in eastern Texas to south-central Oklahoma.
Subspecies brachycarpa was published initially by Peng as a new combination based on Ludwigia cylindrica var. brachycarpa Torrey & A. Gray, not realizing that the variety was based on Jussiaea brachycarpa Lamarck, which Peng considered to be a synonym of L. glandulosa subsp. glandulosa. Therefore, subsp. brachycarpa is a new subspecies but was invalid when published by Peng since it was not accompanied by a Latin description in 1986; that situation has since been remedied.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genus 1, species 82 (31 in the flora): North America, Mexico, West Indies, Bermuda, Central America, South America, se Asia, Africa, Indian Ocean Islands, Pacific Islands, Australasia; introduced in Europe, w Asia.
Ludwigioideae were segregated as a distinct subfamily (W. L. Wagner et al. 2007) to reflect the phylogenetic relationship of Ludwigia as sister to other genera of Onagraceae in morphological and molecular analyses (see R. A. Levin et al. 2003, 2004). Ludwigia is distinguished by the absence of a floral tube, persistence of sepals on capsules after other floral parts dehisce, pollen shed in tetrads or polyads (or as monads in some sections, tetrads sometimes found elsewhere in Onagraceae), double ovule vascular supply, uniquely including a central supply (R. H. Eyde 1981), single-celled ovule archesporium (H. Tobe and P. H. Raven 1996), and a base chromosome number of x = 8.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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