Ludwigia curtissii |
Ludwigia alternifolia |
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Curtiss' primrose-willow |
bushy seedbox, ludwigie à feuilles alternes, rattlebox, seedbox, square-pod water-primrose |
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Habit | Herbs rarely creeping and rooting at nodes, stolons usually absent. | |
Roots | fusiform, fascicled, thickened, epidermis splitting or peeling near base. |
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Stems | erect or ascending at base, very rarely prostrate, unbranched to well branched, branches sometimes very slender, 15–75 cm, glabrous, with slightly raised lines decurrent from leaf axils. |
subterete or somewhat angled, with narrow raised lines or wings decurrent from leaf axils, 40–150 cm, well branched in distal 1/2, glabrous or sparsely to densely strigillose. |
Leaves | alternate; stipules reddish purple, narrowly ovate, 0.2–0.3 × 0.1–0.3 mm, succulent; petiole winged, 0.3–1.2 cm, blade usually oblanceolate-spatulate to spatulate or oblanceolate, rarely sublinear, 1–2.5(–3) × 0.1–0.8 cm, base attenuate, margins subentire with hydathodal glands, apex acute or mucronate, surfaces glabrous; bracts not reduced. |
stipules narrowly deltate, 0.05–0.2 × 0.05–0.1 mm; petiole 0.1–0.3(–0.7) cm; blade lanceolate-elliptic, (0.6–)2–12 × (0.3–)1–1.5(–2.5) cm, base attenuate, margins entire, apex acute, surfaces strigillose throughout or glabrate with strigillose veins; bracts often reduced and more linear. |
Inflorescences | usually not congested, leafy racemes or spikes, flowers solitary in leaf axils; bracteoles attached in opposite pairs near base of ovary, narrowly lanceolate, lanceolate-elliptic, or oblong-linear, 1.5–3.5(–4) × 0.4–0.8 mm, swollen at base, apex acuminate. |
leafy racemes, flowers solitary in leaf axils; bracteoles linear-lanceolate, 1–2.5 mm, margins entire, apex acute or subacuminate, glabrous or with scattered hairs, attached near base of ovary. |
Flowers | sepals ascending, green fading to white near base, ovate-deltate, 1.5–3 × 1.2–2 mm, margins entire, apex narrowly acute or acuminate, surfaces glabrous; petals 0(–3), narrowly elliptic or spatulate, 1–2.5 × 0.5–1 mm, base attenuate, apex obtuse; filaments pale yellow, 0.8–1(–1.3) mm, anthers 0.3–0.6 × 0.3–0.5 mm; pollen shed singly; ovary obovate-obpyramidal, 2–2.5 × 1.8–2.3 mm, glabrous; nectary disc elevated 0.3–0.4 mm on ovary apex, green, 0.9–1.6 mm diam., prominently 4-lobed, glabrous; style pale green, 0.4–0.7 mm, glabrous, stigma pale yellow, subglobose, 0.3–0.4 × 0.2–0.4 mm, not exserted beyond anthers. |
sepals narrowly ovate-deltate, (6–)6.5–9.5 × 4–6.5 mm, margins entire, apex acute to obtuse, surfaces strigillose, sometimes mixed with villous hairs, or glabrate; petals cordate, 10–14 ×8–12 mm, base attenuate, apex emarginate; filaments opaque white, awl-shaped, 1–3 mm, anthers 1–1.7 × 0.6–0.8 mm; ovary subcuboid to globose, 2.5–4 ×2.5–3.5 mm; nectary disc slightly elevated on ovary apex, 0.8–1.5 mm diam., 4-lobed, ringed with soft, curly hairs or glabrous; style 1.5–2.4 mm, glabrous, stigma light yellow, capitate to hemispherical, 1–1.3 × 1.4–2 mm, shallowly 4-lobed, not or scarcely exserted beyond anthers. |
Capsules | obconic, obscurely 4-angled, (2–)2.5–4(–4.7) × 2–3(–3.5) mm, hard-walled, dehiscent by loculicidal slits, pedicel 0.1–0.5 mm. |
subcuboid to squarish globose, 4–6(–7) × 4–6 mm, 4-angled and 4-winged, wings 0.3–1.5 mm wide, pedicel 2–7 mm. |
Seeds | light brown, ellipsoid, 0.4–0.6 × 0.3–0.4 mm, surface cells transversely elongate, glabrous or, sometimes, with surface wax that mimics appressed hairs. |
light brown, oblong to reniform, 0.5–0.8 × 0.2–0.4 mm, surface cells elongate transversely to seed length. |
2n | = 64. |
= 16. |
Ludwigia curtissii |
Ludwigia alternifolia |
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Phenology | Flowering Mar–Nov (year-round). | Flowering summer–early fall. |
Habitat | Pine savannas and flatwoods, marshes, edges of ponds and streams, sandy or peaty swales, limestone prairies, solution pits on limestone. | Swamps, damp, peaty places, roadside ditches, margins of cultivated fields. |
Elevation | 0–50 m. (0–200 ft.) | 0–1800 m. (0–5900 ft.) |
Distribution |
FL; West Indies (Bahamas) |
AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON; QC
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Discussion | Ludwigia curtissii is the only octoploid (n = 32) in sect. Isnardia, and is restricted to peninsular Florida and the Bahamas. It and L. simpsonii they are the only members of the section that do not form true stolons; instead they simply sprout new shoots from the base. The two species are similar in many ways and appear to share three genomes (C. I. Peng 1988). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ludwigia alternifolia is widespread and common in the eastern half of the flora area, as far west as Ontario in Canada, eastern Colorado, and Texas. Ludwigia macrocarpa Michaux 1803 is a superfluous, illegitimate name for L. alternifolia. Rhexia linearifolia Poiret was originally described as a species of Melastomataceae, but the description is superficially similar to Ludwigia and has often been included in synonymy with L. alternifolia. The type of R. linearifolia has not been located or studied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | L. spathulifolia | Isnardia alternifolia, I. alternifolia var. salicifolia, I. alternifolia var. uniflora, I. aurantiaca, L. alternifolia var. linearifolia, L. alternifolia var. pubescens, L. angustifolia var. ramosissima, L. aurantiaca, L. ramosissima, L. salicifolia, L. uniflora |
Name authority | Chapman: Fl. South. U.S. ed. 2, 621. (1883) | Linnaeus: Sp. Pl. 1: 118. (1753) |
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