The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

Carolina primrose-willow

bushy seedbox, ludwigie à feuilles alternes, rattlebox, seedbox, square-pod water-primrose

Habit Herbs, from woody rootstock.
Roots

fusiform, fascicled, thickened, epidermis splitting or peeling near base.

Stems

erect, subterete, 20–120 cm, branched, glabrate proximally, or strigillose, especially in distal parts, with raised strigillose lines decurrent from leaf axils mid stem.

subterete or somewhat angled, with narrow raised lines or wings decurrent from leaf axils, 40–150 cm, well branched in distal 1/2, glabrous or sparsely to densely strigillose.

Leaves

stipules narrowly deltate, 0.3–0.5 × 0.1–0.2 mm;

petiole winged, 0.1–0.7 cm;

blade narrowly to broadly lanceolate, 4–15 × 0.3–1(–3) cm, base tapered, margins subentire to inconspicuously glandular-serrulate, apex acute to acuminate, surfaces finely strigillose, especially on abaxial veins, sometimes glabrate;

bracts narrower, reduced in size.

stipules narrowly deltate, 0.05–0.2 × 0.05–0.1 mm;

petiole 0.1–0.3(–0.7) cm;

blade lanceolate-elliptic, (0.6–)2–12 × (0.3–)1–1.5(–2.5) cm, base attenuate, margins entire, apex acute, surfaces strigillose throughout or glabrate with strigillose veins;

bracts often reduced and more linear.

Inflorescences

open, leafy racemes, flowers solitary in axils;

bracteoles lanceolate-linear or setaceous, 2–6 × 0.3–0.8 mm, attached on pedicel just proximal to base of ovary.

leafy racemes, flowers solitary in leaf axils;

bracteoles linear-lanceolate, 1–2.5 mm, margins entire, apex acute or subacuminate, glabrous or with scattered hairs, attached near base of ovary.

Flowers

sepals ovate-deltate, 10–20 × 7–12 mm, abruptly acuminate or acute, inconspicuously 5–7-nerved, surfaces strigillose;

petals deep golden yellow, broadly obovate, 20–35 × 10–30 mm, apex shallowly emarginate, claw 1.5–3 mm;

stamens 8 in 2 unequal series, filaments flattened and dilated near base, epipetalous set 3.4–4.5 mm, episepalous set 4.5–5.5 mm, anthers oblong, 4–5 mm;

pollen shed in polyads;

ovary subcylindric, slightly 4-angled, 8–12(–20) mm;

nectary disc slightly elevated on ovary apex, 2–3 mm diam., 4-lobed, ringed by short hairs;

style 3–3.5 mm, stigma clavate-capitate, 2.5–3 × 2–2.5 mm, often exserted beyond anthers.

sepals narrowly ovate-deltate, (6–)6.5–9.5 × 4–6.5 mm, margins entire, apex acute to obtuse, surfaces strigillose, sometimes mixed with villous hairs, or glabrate;

petals cordate, 10–14 ×8–12 mm, base attenuate, apex emarginate;

filaments opaque white, awl-shaped, 1–3 mm, anthers 1–1.7 × 0.6–0.8 mm;

ovary subcuboid to globose, 2.5–4 ×2.5–3.5 mm;

nectary disc slightly elevated on ovary apex, 0.8–1.5 mm diam., 4-lobed, ringed with soft, curly hairs or glabrous;

style 1.5–2.4 mm, glabrous, stigma light yellow, capitate to hemispherical, 1–1.3 × 1.4–2 mm, shallowly 4-lobed, not or scarcely exserted beyond anthers.

Capsules

clavate-cylindric, subterete to obtusely 4-angled, 20–35 × 3.5–5 mm, thin walls, irregularly dehiscent, tapering to pedicel 10–40 mm.

subcuboid to squarish globose, 4–6(–7) × 4–6 mm, 4-angled and 4-winged, wings 0.3–1.5 mm wide, pedicel 2–7 mm.

Seeds

in several indistinct rows per locule, yellow-brown, oblong (appearing round), 0.5 mm, shiny, raphe 2/3 as wide as body.

light brown, oblong to reniform, 0.5–0.8 × 0.2–0.4 mm, surface cells elongate transversely to seed length.

2n

= 16.

= 16.

Ludwigia bonariensis

Ludwigia alternifolia

Phenology Flowering summer. Flowering summer–early fall.
Habitat Wet places, mainly along coastal areas, especially ditches, banks near brackish water. Swamps, damp, peaty places, roadside ditches, margins of cultivated fields.
Elevation 0–200[–2600] m. (0–700[–8500] ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
AL; FL; MS; NC; SC; TX; Mexico (Chihuahua, Durango, Puebla, Quintana Roo, Tabasco, Tamaulipas, Veracruz); South America (Argentina, Bolivia, Brazil, Paraguay, Uruguay)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Ludwigia alternifolia is widespread and common in the eastern half of the flora area, as far west as Ontario in Canada, eastern Colorado, and Texas.

Ludwigia macrocarpa Michaux 1803 is a superfluous, illegitimate name for L. alternifolia. Rhexia linearifolia Poiret was originally described as a species of Melastomataceae, but the description is superficially similar to Ludwigia and has often been included in synonymy with L. alternifolia. The type of R. linearifolia has not been located or studied.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Macrocarpon Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Ludwigia
Sibling taxa
L. alata, L. alternifolia, L. arcuata, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
L. alata, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
Synonyms Jussiaea bonariensis, J. neglecta, J. suffruticosa var. bonariensis Isnardia alternifolia, I. alternifolia var. salicifolia, I. alternifolia var. uniflora, I. aurantiaca, L. alternifolia var. linearifolia, L. alternifolia var. pubescens, L. angustifolia var. ramosissima, L. aurantiaca, L. ramosissima, L. salicifolia, L. uniflora
Name authority (Micheli) H. Hara: J. Jap. Bot. 28: 291. (1953) Linnaeus: Sp. Pl. 1: 118. (1753)
Web links