Lomatium minus |
Lomatium planosum |
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Day Valley desert-parsley, John Day desert parsley, John Day Valley desert parsley |
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Habit | Herbs blue-green, acaulous or short-caulescent, 10–30 cm, robust, glabrous; caudex simple or 2–3-branched, with persistent leaf sheaths weathering into fibrous thatch, with persistent, gray peduncles; taproot thick, sometimes horizontal, sometimes with shallow, irregular, tuberlike swellings. | Herbs blue-green or gray-green, acaulous, 10–30 cm, glabrous or scabrous to papillate; caudex branched, usually with persistent leaf sheaths weathering to short, chaffy or chartaceous scales, without persistent peduncles; taproot slender. |
Leaves | arising at slightly different heights, not forming just 1 rosette, blue-green, glaucous, often 2–3-ternate-3-pinnately dissected; petiole broadly sheathing basally to 1/2 length; blade triangular to ovate, 5–12 × 2.7–10 cm, surfaces glabrous; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 1000–5000, linear, 1–5 × 0.5 mm, not overlapping, margins entire, apex acute, callus tips 0–0.2 mm, firm but not spinelike, terminal segment 1–5 mm; cauline leaves 0–2, petioles sometimes sheathing more than 1/2 length. |
forming 1 rosette of 4–20 leaves atop pseudoscape, ± bluish green, ternate-1-pinnate-1-pinnatifid; petiole usually sheathing basally, rarely 1/2 or entire length; blade oblong or lanceolate, 1–6 × 1–3 cm, surfaces glabrous, scabrous, or papillate; ultimate segments 50–400, ovate, 1–5(–8) × 0.5–1.3 mm, margins entire, apex acute, callus tips 0–0.1 mm, terminal segment 0.5–1.5(–3) mm; cauline leaves 0. |
Pseudoscapes | absent or subterranean. |
present, often aerial, 2–15 cm. |
Peduncles | 1–6 per plant, usually 1 per stem, decumbent, spreading, or ascending, strongly inflated at maturity, 5–15(–24) cm, exceeding leaves, 2–8(–11) mm wide 1 cm below umbel, glabrous. |
1–20 per plant, 1 per stem, ascending to erect, not inflated, 2–20 cm, exceeding leaves, 1(–2) mm wide 1 cm below umbel, glabrous, scabrous, or papillate. |
Umbels | 2.5–4.7 cm wide in flower, 3.6–8.6 cm wide in fruit, rays 6–16, spreading, 1–4(–6) cm in fruit, subequal to unequal, glabrous; involucel bractlets several, distinct, linear-subulate, (3–)4–9(–15) mm, shorter or longer than flowers, margins very broadly scarious, not ciliate, entire, glabrous; umbellets 8–15-flowered. |
0.8–2.5 cm wide in flower, 3.5–10 cm wide in fruit, rays 3–13, ascending, then broadly spreading or sometimes reflexed in fruit, 1.5–3(–4.5) cm in fruit, unequal, glabrous or papillate to scabrous on veins; involucel bractlets 3–5, usually distinct, linear, 1–3(–4) mm, those subtending pedicels of male flowers usually longer than those subtending pedicels of bisexual flowers, subequal to flowers, margins scarious or not, not ciliate or shortly so, entire, glabrous. |
Flowers | petals purple to dark pink, glabrous; anthers purple; ovary and young fruit glabrous. |
petals yellow, rarely purple; glabrous; anthers yellow to purple; ovary and young fruit glabrous. |
Fruiting pedicels | (5.5–)6.5–8(–9) mm, shorter than fruit. |
2–5 mm, shorter than fruit. |
Mericarps | ± dorsiventrally compressed, narrowly elliptic or oblong-oval, 8.8–16(–19.3) × (3–)4.7–7.8 mm, length/width ratio 1.9–3.3; wings 0.9–2 mm wide, 25–50% of body width, ± same color as body; abaxial ribs slightly raised; apex obtuse; oil ducts usually 1 in intervals, 3–4 on commissure, conspicuous. |
not dorsiventrally compressed, broadly oblong, 5–7 × 2.2–3.1 mm, length/width ratio 2–2.6; lateral wing 0.4–0.8 mm wide, 33–55% of body width, paler than body; abaxial ribs prominately raised, forming wings 0.3–0.4 mm wide, about as high as lateral wings; apex obtuse to rounded; oil ducts 3–4 in intervals, 4–7 on commissure. |
Lomatium minus |
Lomatium planosum |
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Phenology | Flowering (Mar–)Apr–May; fruiting May–Jun. | Flowering Apr–May; fruiting May–Jun. |
Habitat | Steep, unstable talus slopes, stone stripes, rock outcrops. | Meadows, slopes, riparian areas, ridgetops, sagebrush scrub, pinyon-juniper woodlands, ponderosa pine woodlands, Douglas-fir forests, Englemann spruce forests, oak woodlands, barren sites, usually on shale, sometimes on limestone, sandstone, or clay substrates. |
Elevation | (700–)1000–1300 m. [(2300–)3300–4300 ft.] | 1700–2800 m. [5600–9200 ft.] |
Distribution |
OR
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CO |
Discussion | Lomatium minus is strongly glaucous with purple or pink petals, narrow leaflets, and an inflated stem like that of L. columbianum. However, L. minus is a much smaller plant, and the peduncle is inflated unevenly. In mature fruits, the wings curve back, making each mericarp rounded in cross section like a bread roll. Lomatium minus is endemic to the Blue Mountains region of central Oregon, with an outlying population in northern Malheur County. It is sometimes confused with L. tuberosum, which has similar petal colors and leaflets but is endemic to central Washington. Lomatium minus is a culturally significant food plant to members of the Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lomatium planosum is known from Delta, Eagle, Garfield, Mesa, Moffat, Montrose, Pitkin, Rio Blanco, and Routt counties. It was long classified in Cymopterus because its abaxial ribs are raised, forming thick wings. However, DNA-based analysis shows that it is more closely related to Lomatium species than to Cymopterus (D. H. Mansfield et. al 2017). It has an unusual growth form. The pseudoscape usually extends above ground, where it produces a rosette of leaves. Lomatium planosum leaves are like those of C. longipes, but that species has yellow or white petals, fruiting pedicels 3–8 mm, and lateral fruit wings 1–1.5(–2) mm wide, and is not known from Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 13. | FNA vol. 13. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Leptotaenia minor | Aulospermum planosum, Cymopterus planosus |
Name authority | (Rose ex Howell) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) | (Osterhout) Mansfield & S. R. Downie: Phytotaxa 316: 97. (2017) |
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