Lomatium minus |
Lomatium piperi |
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Day Valley desert-parsley, John Day desert parsley, John Day Valley desert parsley |
Indian biscuitroot, Piper's biscuit-root, Piper's desert-parsley, Piper's lomatium, salt-and-pepper |
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Habit | Herbs blue-green, acaulous or short-caulescent, 10–30 cm, robust, glabrous; caudex simple or 2–3-branched, with persistent leaf sheaths weathering into fibrous thatch, with persistent, gray peduncles; taproot thick, sometimes horizontal, sometimes with shallow, irregular, tuberlike swellings. | Herbs green to slightly blue-green, caulescent, 7–20(–25) cm, relatively delicate, glabrous or hairy; caudex simple, with or without persistent leaf sheaths weathering into sparse thatch of fibers and chaffy or chartaceous scales, without persistent peduncles; taproot with shallow or deep, globose or ovoid, tuberlike swellings. |
Leaves | arising at slightly different heights, not forming just 1 rosette, blue-green, glaucous, often 2–3-ternate-3-pinnately dissected; petiole broadly sheathing basally to 1/2 length; blade triangular to ovate, 5–12 × 2.7–10 cm, surfaces glabrous; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 1000–5000, linear, 1–5 × 0.5 mm, not overlapping, margins entire, apex acute, callus tips 0–0.2 mm, firm but not spinelike, terminal segment 1–5 mm; cauline leaves 0–2, petioles sometimes sheathing more than 1/2 length. |
arising at slightly different heights, not forming just 1 rosette, green, 1–2(–3)-ternate-(0–)1–2-pinnate-0–2-pinnatifid, basal leaves with petiole sheathing basally; blade triangular to ovate, 3–7.5 × 1.5–5.5 cm, surfaces glabrous; leaflets not overlapping, penultimate segments narrow, usually less than 2 mm wide, ultimate segments 10–80, linear, 2–30(–40) × 0.5–2 mm, larger ones longer than 6 mm, margins entire, usually not reflexed, apex acute to obtuse or rounded, callus tips 0–0.3 mm, firm but not spinelike, terminal segment 3–20 mm; cauline leaves 1(–2), distal ones much reduced with 1–3 ultimate segments. |
Pseudoscapes | absent or subterranean. |
subterranean. |
Peduncles | 1–6 per plant, usually 1 per stem, decumbent, spreading, or ascending, strongly inflated at maturity, 5–15(–24) cm, exceeding leaves, 2–8(–11) mm wide 1 cm below umbel, glabrous. |
1–3 per plant, 1–2 per stem, decumbent to erect, not inflated, 2–8 cm, exceeding leaves, 0.5–2.5 mm wide 1 cm below umbel, glabrous. |
Umbels | 2.5–4.7 cm wide in flower, 3.6–8.6 cm wide in fruit, rays 6–16, spreading, 1–4(–6) cm in fruit, subequal to unequal, glabrous; involucel bractlets several, distinct, linear-subulate, (3–)4–9(–15) mm, shorter or longer than flowers, margins very broadly scarious, not ciliate, entire, glabrous; umbellets 8–15-flowered. |
0.6–3.1 cm wide in flower, 1.5–4.4 cm and ± congested in fruit, rays 3–20, spreading, (1–)1.5–3.5(–5) cm in fruit, unequal, glabrous or sparsely scaberulous; involucel bractlets (0 or)2–6, distinct, linear or lanceolate, 0.5–2.2 mm, shorter than flowers, margins broadly scarious, not ciliate, entire, glabrous. |
Flowers | petals purple to dark pink, glabrous; anthers purple; ovary and young fruit glabrous. |
petals white, glabrous; anthers purple; ovary and young fruit glabrous. |
Fruiting pedicels | (5.5–)6.5–8(–9) mm, shorter than fruit. |
0.5–2.5 mm, shorter than fruit, spreading to erect when fruit is mature. |
Mericarps | ± dorsiventrally compressed, narrowly elliptic or oblong-oval, 8.8–16(–19.3) × (3–)4.7–7.8 mm, length/width ratio 1.9–3.3; wings 0.9–2 mm wide, 25–50% of body width, ± same color as body; abaxial ribs slightly raised; apex obtuse; oil ducts usually 1 in intervals, 3–4 on commissure, conspicuous. |
dorsiventrally compressed, oblong, broadly elliptic, elliptic, or ovate, 4–9 × 1.9–4.5 mm, length/width ratio 1.4–2.5; wings 0.7–1.5 mm wide, 40–70% of body width, paler than body, well developed, not corky-thickened, spreading, flat; abaxial ribs slightly raised; apex obtuse to rounded; oil ducts 1–8 in intervals, 2–7 on commissure. |
2n | = 22. |
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Lomatium minus |
Lomatium piperi |
|
Phenology | Flowering (Mar–)Apr–May; fruiting May–Jun. | Flowering late Feb–mid Jul; fruiting Mar–Jul. |
Habitat | Steep, unstable talus slopes, stone stripes, rock outcrops. | Mesic sagebrush-bunchgrass steppe, edges of ponderosa pine forests, rocky slopes and flats in scablands. |
Elevation | (700–)1000–1300 m. [(2300–)3300–4300 ft.] | 50–1700 m. [160–5600 ft.] |
Distribution |
OR
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CA; ID; OR; WA
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Discussion | Lomatium minus is strongly glaucous with purple or pink petals, narrow leaflets, and an inflated stem like that of L. columbianum. However, L. minus is a much smaller plant, and the peduncle is inflated unevenly. In mature fruits, the wings curve back, making each mericarp rounded in cross section like a bread roll. Lomatium minus is endemic to the Blue Mountains region of central Oregon, with an outlying population in northern Malheur County. It is sometimes confused with L. tuberosum, which has similar petal colors and leaflets but is endemic to central Washington. Lomatium minus is a culturally significant food plant to members of the Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
White petals and purple anthers give tiny Lomatium piperi the common name of pepper-and-salt, a name shared with the similar L. geyeri and L. gormanii. Lomatium piperi plants are usually caulescent, unlike the similar species, although sometimes the only evidence of that is a reduced, bractlike, cauline leaf. In addition, roots of L. geyeri usually have 2+ tuberlike swellings, whereas L. piperi and L. gormanii have one. Ovaries and young fruits of L. gormanii are roughened with as many as 200 papillae per mm2, whereas in L. piperi and L. geyeri, they are glabrous or occasionally sparsely roughened with up to 5 papillae per mm2. Fruits of all three species are usually glabrous when fully mature. Flowering L. linearifolium is nearly impossible to distinguish from flowering L. piperi, unless the habitat and location are known. In fruit, they are easily distinguished because the mature fruit of L. linearifolium is more or less round in cross section, whereas that of L. piperi is dorsiventrally compressed. Furthermore, the immature fruits of L. piperi have smooth abaxial surfaces, whereas those of L. linearifolium have slightly raised abaxial ribs; this can aid in identification when the fruits are not fully mature. L. piperi grows in usually seasonally wet rocky soils on scablands, basins, and open slopes in California, Idaho, Oregon, and Washington. In contrast, L. linearifolium grows in usually moist sandy or loamy soil near vernal snowbanks, the plants blooming as soon as snow melts. Its range is in Washington and northern Oregon east of the Cascade Range, as well as Idaho, southern Montana, and northwestern Wyoming. Lomatium piperi is a culturally significant food plant for members of the Paiute and Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 13. | FNA vol. 13. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Leptotaenia minor | |
Name authority | (Rose ex Howell) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) | J. M. Coulter & Rose: Contr. U.S. Natl. Herb. 7: 211. (1900) |
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