Lomatium minus |
Lomatium klickitatense |
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Day Valley desert-parsley, John Day desert parsley, John Day Valley desert parsley |
Klickitat biscuit-root, Klickitat desert-parsley, Klickitat lomatium |
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Habit | Herbs blue-green, acaulous or short-caulescent, 10–30 cm, robust, glabrous; caudex simple or 2–3-branched, with persistent leaf sheaths weathering into fibrous thatch, with persistent, gray peduncles; taproot thick, sometimes horizontal, sometimes with shallow, irregular, tuberlike swellings. | Herbs green, caulescent or short-caulescent, rarely acaulous, 40–150 cm, glabrous; caudex simple to multicipital, with persistent leaf sheaths weathering into dense thatch of fibers and chaffy scales, with few persistent, gray peduncles; taproot thick, sometimes with deep, irregular tuberlike swellings. |
Leaves | arising at slightly different heights, not forming just 1 rosette, blue-green, glaucous, often 2–3-ternate-3-pinnately dissected; petiole broadly sheathing basally to 1/2 length; blade triangular to ovate, 5–12 × 2.7–10 cm, surfaces glabrous; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 1000–5000, linear, 1–5 × 0.5 mm, not overlapping, margins entire, apex acute, callus tips 0–0.2 mm, firm but not spinelike, terminal segment 1–5 mm; cauline leaves 0–2, petioles sometimes sheathing more than 1/2 length. |
arising at slightly different heights, not forming just 1 rosette, green, ternate-2–4-pinnate or -pinnatifid; petiole sheathing basally to entire length, usually equaling or shorter than blade; blade ovate, 10–30 × 5–23 cm, surfaces glabrous; apical 2–3 pinnule pairs of secondary leaflets with naked intercostal region between rachilla and basalmost pinnule lobes, appearing like petiolule, this “petiolule” of penultimate segments (1–)2–6 mm; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 300–1500+, narrowly linear, subterete in cross section, (6.5–)7–15 × 0.1–0.4 mm, appearing spread along leaf because naked portions of petiolules are usually shorter than part of leaflet that has blades, diffuse, not obscuring elongate intercostal areas along rachillae (best assessed at arm’s length), margins entire, apex acuminate, callus tips 0–0.1 mm, terminal segment 0.7–1.2 mm; cauline leaves (0–)1–2. |
Pseudoscapes | absent or subterranean. |
absent. |
Peduncles | 1–6 per plant, usually 1 per stem, decumbent, spreading, or ascending, strongly inflated at maturity, 5–15(–24) cm, exceeding leaves, 2–8(–11) mm wide 1 cm below umbel, glabrous. |
1–10+ per plant, 1 per stem, spreading to erect, not inflated, 30–60 cm, slender or stout, equaling leaves in flower, exceeding them in fruit, 3–8 mm wide wide 1 cm below umbel, glabrous. |
Umbels | 2.5–4.7 cm wide in flower, 3.6–8.6 cm wide in fruit, rays 6–16, spreading, 1–4(–6) cm in fruit, subequal to unequal, glabrous; involucel bractlets several, distinct, linear-subulate, (3–)4–9(–15) mm, shorter or longer than flowers, margins very broadly scarious, not ciliate, entire, glabrous; umbellets 8–15-flowered. |
0.7–4.5 cm wide in flower, 12–20 cm wide in fruit, rays 14–30, ascending to spreading, 4–12 cm in fruit, unequal to subequal, glabrous; involucel bractlets (0 or)2–10, distinct, linear to lanceolate, 2–10 mm, shorter than flowers, margins narrowly scarious, not ciliate, entire, glabrous. |
Flowers | petals purple to dark pink, glabrous; anthers purple; ovary and young fruit glabrous. |
petals yellow, glabrous; anthers yellow or ochroleucous; ovary and young fruit glabrous. |
Fruiting pedicels | (5.5–)6.5–8(–9) mm, shorter than fruit. |
6–15 mm, shorter than fruit. |
Mericarps | ± dorsiventrally compressed, narrowly elliptic or oblong-oval, 8.8–16(–19.3) × (3–)4.7–7.8 mm, length/width ratio 1.9–3.3; wings 0.9–2 mm wide, 25–50% of body width, ± same color as body; abaxial ribs slightly raised; apex obtuse; oil ducts usually 1 in intervals, 3–4 on commissure, conspicuous. |
dorsiventrally compressed, oblong or elliptic, (9–)12–18 × 5–8 mm, length/width ratio 2–2.6; wings (0.7–)1–1.4 mm wide, (15–)20–35(–40)% of body width, paler than body; abaxial ribs slightly raised; apex obtuse to narrowly rounded; oil ducts 1–2 in intervals, 2–6 on commissure. |
Lomatium minus |
Lomatium klickitatense |
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Phenology | Flowering (Mar–)Apr–May; fruiting May–Jun. | Flowering Apr–May; fruiting May–Jun. |
Habitat | Steep, unstable talus slopes, stone stripes, rock outcrops. | Rocky hillsides, talus slopes, cliffs, open rocky sites in oak or conifer forests. |
Elevation | (700–)1000–1300 m. [(2300–)3300–4300 ft.] | 100–800 m. [300–2600 ft.] |
Distribution |
OR
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OR; WA
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Discussion | Lomatium minus is strongly glaucous with purple or pink petals, narrow leaflets, and an inflated stem like that of L. columbianum. However, L. minus is a much smaller plant, and the peduncle is inflated unevenly. In mature fruits, the wings curve back, making each mericarp rounded in cross section like a bread roll. Lomatium minus is endemic to the Blue Mountains region of central Oregon, with an outlying population in northern Malheur County. It is sometimes confused with L. tuberosum, which has similar petal colors and leaflets but is endemic to central Washington. Lomatium minus is a culturally significant food plant to members of the Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lomatium klickitatense is endemic to the Columbia River Gorge in Hood River and Wasco counties, Oregon, and Klickitat County, Washington. It produces diffuse, cloudlike mounds of narrowly linear leaflets, much like those of L. columbianum but green rather than glaucous. Also, its petals are yellow, and its fruits are smaller than those of L. columbianum. The two species sometimes grow together. Lomatium klickitatense is similar to L. papilioniferum, which often has blue-green foliage covered with tiny glands, and ultimate leaf segments that are broader, shorter, and rounded. The difference can be obscured by the tendency for L. klickitatense to be collected when it is immature and leaves have not reached their full length, and for L. papilioniferum growing in shade to have somewhat longer and greener ultimate leaf segments. One specimen from Idaho County, Idaho, appears to be L. klickitatense. It is unclear if this represents an established population, an extreme of variation of L. brunsfeldianum, or a waif resulting from trade among the Sahaptin Native nations and the Nez Perce. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 13. | FNA vol. 13. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Leptotaenia minor | |
Name authority | (Rose ex Howell) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) | J. A. Alexander & Whaley: J. Bot. Res. Inst. Texas 12: 408, figs. 1I-J, 9. (2018) |
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