Lomatium minus |
Lomatium grayi |
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Day Valley desert-parsley, John Day desert parsley, John Day Valley desert parsley |
Gray's biscuitroot, Gray's lomatium, pungent desert parsley |
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Habit | Herbs blue-green, acaulous or short-caulescent, 10–30 cm, robust, glabrous; caudex simple or 2–3-branched, with persistent leaf sheaths weathering into fibrous thatch, with persistent, gray peduncles; taproot thick, sometimes horizontal, sometimes with shallow, irregular, tuberlike swellings. | Herbs green or ± blue-green, acaulous, 8–20 cm, glabrous; caudex simple or multicipital, with persistent leaf sheaths weathering into dense thatch of fibers and chaffy scales, without persistent peduncles; taproot thick, sometimes with deep, irregular tuberlike swellings. |
Leaves | arising at slightly different heights, not forming just 1 rosette, blue-green, glaucous, often 2–3-ternate-3-pinnately dissected; petiole broadly sheathing basally to 1/2 length; blade triangular to ovate, 5–12 × 2.7–10 cm, surfaces glabrous; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 1000–5000, linear, 1–5 × 0.5 mm, not overlapping, margins entire, apex acute, callus tips 0–0.2 mm, firm but not spinelike, terminal segment 1–5 mm; cauline leaves 0–2, petioles sometimes sheathing more than 1/2 length. |
arising at slightly different heights, not forming just 1 rosette, green or blue-green, ternate-1–4-pinnate or -pinnatifid; petiole sheathing basally to entire length; blade broadly elliptic to rhombic, 6–12 × 4–7 cm, surfaces glabrous; apical 2–3 pinnule pairs of secondary leaflets without naked intercostal region between rachilla and basalmost pinnule lobes, appearing sessile; ultimate segments 1000–3500, very narrowly linear, subterete in cross section, 0.8–4.8 × 0.1–0.4 mm, congested, obscuring short intercostal areas along rachillae, margins entire, apex acuminate, callus tips 0–0.2 mm, terminal segment 3 mm; cauline leaves 0. |
Pseudoscapes | absent or subterranean. |
often present, subterranean. |
Peduncles | 1–6 per plant, usually 1 per stem, decumbent, spreading, or ascending, strongly inflated at maturity, 5–15(–24) cm, exceeding leaves, 2–8(–11) mm wide 1 cm below umbel, glabrous. |
1–5 per plant, 1 per stem, ascending or erect, not inflated, 8–40 cm, exceeding leaves, 0.6–3.5(–6) mm wide 1 cm below umbel, glabrous. |
Umbels | 2.5–4.7 cm wide in flower, 3.6–8.6 cm wide in fruit, rays 6–16, spreading, 1–4(–6) cm in fruit, subequal to unequal, glabrous; involucel bractlets several, distinct, linear-subulate, (3–)4–9(–15) mm, shorter or longer than flowers, margins very broadly scarious, not ciliate, entire, glabrous; umbellets 8–15-flowered. |
2.5–4 cm wide in flower, 5–13 cm wide in fruit, rays 5–22, erect to ascending, 0.5–8 cm in fruit, subequal (except sterile ones), glabrous; involucel bractlets (0 or)2–10, distinct or connate, sometimes connate in pairs and thus appearing 2-fid, linear or lanceolate, 2–10 mm, subequal to or longer than flowers, margins green or scarious, not ciliate, entire or deeply pinnately lobed, glabrous. |
Flowers | petals purple to dark pink, glabrous; anthers purple; ovary and young fruit glabrous. |
sepals scarious or greenish, to 0.6 mm; petals yellow, glabrous; anthers yellow or ochroleucous; ovary and young fruit glabrous. |
Fruiting pedicels | (5.5–)6.5–8(–9) mm, shorter than fruit. |
3–16 mm, shorter than fruit. |
Mericarps | ± dorsiventrally compressed, narrowly elliptic or oblong-oval, 8.8–16(–19.3) × (3–)4.7–7.8 mm, length/width ratio 1.9–3.3; wings 0.9–2 mm wide, 25–50% of body width, ± same color as body; abaxial ribs slightly raised; apex obtuse; oil ducts usually 1 in intervals, 3–4 on commissure, conspicuous. |
dorsiventrally compressed, oblong or elliptic, (5–)8–14 × 4.5–9 mm, length/width ratio 1.3–2.5; wings 1.1–2.3 mm wide, 30–75% of body width, paler than body, thin, not corky-thickened; abaxial ribs raised; apex acute, rounded, broadly rounded; oil ducts 1–2 in intervals, 2–6 on commissure. |
Lomatium minus |
Lomatium grayi |
|
Phenology | Flowering (Mar–)Apr–May; fruiting May–Jun. | Flowering Apr–May; fruiting late May–Jun. |
Habitat | Steep, unstable talus slopes, stone stripes, rock outcrops. | Sagebrush, pinyon-juniper, mountain brush, ponderosa pine, and Douglas-fir communities. |
Elevation | (700–)1000–1300 m. [(2300–)3300–4300 ft.] | 1400–1800 m. [4600–5900 ft.] |
Distribution |
OR
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CO; ID; NM; UT; WY
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Discussion | Lomatium minus is strongly glaucous with purple or pink petals, narrow leaflets, and an inflated stem like that of L. columbianum. However, L. minus is a much smaller plant, and the peduncle is inflated unevenly. In mature fruits, the wings curve back, making each mericarp rounded in cross section like a bread roll. Lomatium minus is endemic to the Blue Mountains region of central Oregon, with an outlying population in northern Malheur County. It is sometimes confused with L. tuberosum, which has similar petal colors and leaflets but is endemic to central Washington. Lomatium minus is a culturally significant food plant to members of the Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Lomatium grayi has been used for diverse species that have much-dissected leaves with very narrowly linear segments arranged in three dimensions. However, swallowtail butterflies (Papilio species) distinguish among these plants, and botanists followed up this clue to separate traditional L. grayi into several species; compare material with L. depauperatum, L. klickitatense, and L. papilioniferum, as well as longer-fruited L. brunsfeldianum and L. filicinum. Lomatium grayi itself is the easternmost of these species, growing from central Utah and southeastern Idaho eastward to southwestern Wyoming, and western Colorado, and northwestern New Mexico. The most similar species is L. depauperatum, and the two co-occur in central Utah; see discussion under 23. L. depauperatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 13. | FNA vol. 13. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Leptotaenia minor | Peucedanum grayi |
Name authority | (Rose ex Howell) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) | (J. M. Coulter & Rose) J. M. Coulter & Rose: Contr. U.S. Natl. Herb. 7: 229. (1900) |
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