Lomatium minus |
Lomatium dissectum |
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Day Valley desert-parsley, John Day desert parsley, John Day Valley desert parsley |
fern-leaf biscuit-root, fern-leaf desert-parsley, fern-leaf lomatium, sessile-fruit fern-leaf biscuit-root |
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Habit | Herbs blue-green, acaulous or short-caulescent, 10–30 cm, robust, glabrous; caudex simple or 2–3-branched, with persistent leaf sheaths weathering into fibrous thatch, with persistent, gray peduncles; taproot thick, sometimes horizontal, sometimes with shallow, irregular, tuberlike swellings. | Herbs green or sometimes slightly glaucous, caulescent, 30–130(–220) cm, robust, apparently glabrous but usually scabrous to hairy with short triangular hairs on leaf margins and veins; caudex simple, 2–3-branched, or multicipital, with persistent leaf sheaths weathering into fibrous thatch at or below ground level, with or without persistent peduncles; taproot slender or irregularly thickened, often massive, lacking distinct tuberlike swellings. |
Leaves | arising at slightly different heights, not forming just 1 rosette, blue-green, glaucous, often 2–3-ternate-3-pinnately dissected; petiole broadly sheathing basally to 1/2 length; blade triangular to ovate, 5–12 × 2.7–10 cm, surfaces glabrous; penultimate segments narrow, usually less than 2 mm wide, ultimate segments 1000–5000, linear, 1–5 × 0.5 mm, not overlapping, margins entire, apex acute, callus tips 0–0.2 mm, firm but not spinelike, terminal segment 1–5 mm; cauline leaves 0–2, petioles sometimes sheathing more than 1/2 length. |
arising at slightly different heights, not forming just 1 rosette, usually green, to slightly blue-green, ternate-pinnately decompound; petiole sheathing basally to entire length; blade ovate to orbiculate, 10–30 × 6–30 cm wide, abaxial surface sparsely to densely scabrous to puberulent with triangular hairs on margins and veins, adaxial surface glabrous, sometimes scabrous to puberulent; ultimate segments 1000–8000, ovate, 1–12 × 0.4–2.5(–4.5) mm, relatively narrow, margins entire, usually not reflexed, apex rounded to acute, callus tips 0–0.1 mm, firm but not spinelike, terminal segment 1–5 mm or 15–30 mm if counting entire area with confluent leaf tissue; cauline leaves 1–2+, usually with more than 5 leaflets. |
Pseudoscapes | absent or subterranean. |
absent. |
Peduncles | 1–6 per plant, usually 1 per stem, decumbent, spreading, or ascending, strongly inflated at maturity, 5–15(–24) cm, exceeding leaves, 2–8(–11) mm wide 1 cm below umbel, glabrous. |
1–25+ per plant, 1–3 per stem, ascending to erect, often glaucous, not inflated, 20–80 cm, exceeding leaves, (2–)3–6 mm wide 1 cm below umbel, not brittle, glabrous. |
Umbels | 2.5–4.7 cm wide in flower, 3.6–8.6 cm wide in fruit, rays 6–16, spreading, 1–4(–6) cm in fruit, subequal to unequal, glabrous; involucel bractlets several, distinct, linear-subulate, (3–)4–9(–15) mm, shorter or longer than flowers, margins very broadly scarious, not ciliate, entire, glabrous; umbellets 8–15-flowered. |
2–16 cm wide in flower, 7–25 cm in fruit, involucral bracts usually present, often fused basally, sometimes fused nearly throughout in 2–3 groups, to 3 cm, rays (9–)15–35, fertile rays 10–30, ascending to spreading, (1–)3–12 cm in fruit, unequal, glabrous; involucel bractlets 3–7, distinct or connate basally, linear to lanceolate, 3–6 mm, or occasionally much longer and leaflike, usually subequal to flowers, sometimes longer, margins narrowly scarious, not ciliate, entire, glabrous. |
Flowers | petals purple to dark pink, glabrous; anthers purple; ovary and young fruit glabrous. |
petals usually brown, maroon-red or purplish, occasionally ± tinged yellow or yellow-green, glabrous; anthers yellow, yellow tinged with brown, yellow-green, or purplish; ovary and young fruit glabrous. |
Fruiting pedicels | (5.5–)6.5–8(–9) mm, shorter than fruit. |
0.6–3(–4) mm, shorter than fruit, shorter than pedicels of staminate flowers. |
Mericarps | ± dorsiventrally compressed, narrowly elliptic or oblong-oval, 8.8–16(–19.3) × (3–)4.7–7.8 mm, length/width ratio 1.9–3.3; wings 0.9–2 mm wide, 25–50% of body width, ± same color as body; abaxial ribs slightly raised; apex obtuse; oil ducts usually 1 in intervals, 3–4 on commissure, conspicuous. |
dorsiventrally compressed, broadly elliptic to oblong, 7–15(–20) × (5.5–)6–8 mm, length/width ratio 1.6–3; wings (0.6–)0.8–1.2 mm wide, 10–30% of body width, same color as or paler than body; abaxial ribs slightly or not raised; apex usually obtuse to rounded; oil ducts obscure. |
2n | = 22. |
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Lomatium minus |
Lomatium dissectum |
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Phenology | Flowering (Mar–)Apr–May; fruiting May–Jun. | Flowering late Mar–May; fruiting May–Jul(–Aug). |
Habitat | Steep, unstable talus slopes, stone stripes, rock outcrops. | Gravelly or rocky slopes, rock outcrops, and open flats in scrubland, woodland, oak or conifer forests, in sagebrush, pinyon-juniper, oak-maple, aspen-fir, riparian, rarely greasewood-desert shrub communities. |
Elevation | (700–)1000–1300 m. [(2300–)3300–4300 ft.] | 150–2000 m. [500–6600 ft.] |
Distribution |
OR
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CA; ID; OR; WA; BC
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Discussion | Lomatium minus is strongly glaucous with purple or pink petals, narrow leaflets, and an inflated stem like that of L. columbianum. However, L. minus is a much smaller plant, and the peduncle is inflated unevenly. In mature fruits, the wings curve back, making each mericarp rounded in cross section like a bread roll. Lomatium minus is endemic to the Blue Mountains region of central Oregon, with an outlying population in northern Malheur County. It is sometimes confused with L. tuberosum, which has similar petal colors and leaflets but is endemic to central Washington. Lomatium minus is a culturally significant food plant to members of the Sahaptin Native nations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lomatium dissectum and its close relative L. multifidum are relatively tall species with very large, stout roots. They are variable, and recognizing two species in this complex may be conservative. Lomatium dissectum occurs mainly west of the Cascade Range and Sierra Nevada, although there are disjunct populations in northern Idaho. Lomatium dissectum usually has brownish petals, giving it the name chocolate tips. Petals may be more or less purple or have a yellowish tint. It is also characterized by short pedicels and glaucous peduncles. Lomatium multifidum usually has yellow petals, although they may be purplish or brown, and has longer pedicels and nonglaucous peduncles. In some L. multifidum populations, leaves are similar to those of L. dissectum, but in others, the leaves are much more finely divided. These two species are sometimes confused with Ligusticum species, which tend to be equally large but have white petals and often more umbels per stem. Lomatium dissectum is a culturally significant food and medicinal plant to members of the Okanagan and Thompson Native nations (D. E. Moerman 1998). See L. multifidum for issues related to exploitation and conservation. Ferula dissecta (Nuttall) A. Gray (1868, not Ledebour 1844) is an illegitimate name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 13. | FNA vol. 13. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Leptotaenia minor | Leptotaenia dissecta |
Name authority | (Rose ex Howell) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) | (Nuttall) Mathias & Constance: Bull. Torrey Bot. Club 69: 246. (1942) |
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