Lithophragma trifoliatum |
Lithophragma |
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fringecup, lithophragma, prairiestar, woodland-star |
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Habit | Plants slender. | Herbs, rhizomatous, not stoloniferous; caudex sometimes bearing bulbils. | ||||||||||||||||||||||||||||||||||||
Flowering stems | simple, 20–50 cm. |
erect, leafy, 8–85 cm, stipitate-glandular. |
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Leaves | in basal rosette and cauline, basal digitately 3-lobed, segments again lobed, cauline (2–3), 3-foliolate or deeply lobed, reduced, similar to basal (except lobes longer); stipules large, decurrent on petiole base; petiole to 11 cm; blade dark green or reddish green, orbiculate, (base hastate), surfaces densely hairy. |
in basal rosette and cauline; cauline leaves (1–)2(–10), usually alternate (opposite in L. cymbalaria), unlobed, lobed, or palmately compound, sometimes with bulbils in axils; stipules present; petiole stipitate-glandular; blade rhombic or orbiculate to reniform, lobed, base cordate, hastate, rounded, ligulate, or cuneate, ultimate margins entire or toothed, apex obtuse or acute, surfaces often stipitate-glandular; venation palmate. |
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Inflorescences | usually 1, (lax), nodding, 4–8-flowered racemes, simple. |
compact or lax racemes, arising from axillary buds in basal rosette, 2–12(–25)-flowered, sometimes flowers solitary, (flowers sometimes replaced by bulbils), bracteate. |
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Pedicels | not exceeding length of hypanthium. |
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Flowers | deciduous if unfertilized, fragrant, horizontal; hypanthium elongate-obconic, elongating in fruit, abruptly expanding, open at throat, (9–11 mm, length 3–4 times diam.); sepals erect, triangular; petals widely spreading, pink, obovate-rhombic, narrowly clawed, 3-lobed, 4–14 mm, ultimate margins entire; ovary 1/2+ to ± entirely inferior; styles included in fruit; stigma papillae in narrow subapical band. |
hypanthium partially adnate to proximal 1/2 to entire length of ovary, free from ovary 1–2 mm, green; sepals 5, green tinged with red; petals 5, white or pink; nectariferous tissue inconspicuous; stamens 10, (inserted on hypanthium, inner whorl attached proximal to point of petal insertion); filaments filiform; (anthers cordate); pistil 3-carpellate; ovary superior to 1/2 or almost completely inferior, 1-locular, carpels fully connate at ovaries; placentation parietal; styles 3; stigmas 3. |
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Capsules | 3-beaked. |
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Seeds | 0.6–0.7 mm, smooth or wrinkled. |
(50–200), usually dark brown, ovoid, smooth, wrinkled, or tuberculate. |
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x | = 7. |
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2n | = 28. |
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Lithophragma trifoliatum |
Lithophragma |
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Phenology | Flowering Mar–May. | |||||||||||||||||||||||||||||||||||||
Habitat | Exposed runnels or near small streams, oak-coniferous woodland | |||||||||||||||||||||||||||||||||||||
Elevation | 40-700 m (100-2300 ft) | |||||||||||||||||||||||||||||||||||||
Distribution |
CA |
w North America |
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Discussion | Lithophragma trifoliatum is known from the western slope of the Cascade Range and Sierra Nevada in California. It is closely related to L. parviflorum and was considered a variety by P. E. Elvander (1993). The shape of the hypanthium, the fragrant flowers, the relatively large pink petals, and the relatively large seeds are distinctive. Lithophragma trifoliatum rarely produces seed; in cultivation, self-pollination was unsuccessful (R. L. Taylor 1965). It may represent a sterile derivative of L. parviflorum that now persists by vegetative reproduction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tellima R. Brown [unranked] Lithophragma Nuttall, J. Acad. Nat. Sci. Philadelphia 7: 26. 1834 (as Lithofragma); Pleurendotria Rafinesque, name rejected Species 10 (10 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 8, p. 80. | FNA vol. 8, p. 77. | ||||||||||||||||||||||||||||||||||||
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Synonyms | L. parviflorum var. trifoliatum | |||||||||||||||||||||||||||||||||||||
Name authority | Eastwood: Bull. Torrey Bot. Club 32: 200. 1905 (as trifoliata), | (Nuttall) Torrey & A. Gray: Fl. N. Amer. 1: 583. 1840, name conserved , | ||||||||||||||||||||||||||||||||||||
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