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fringecup, lithophragma, prairiestar, woodland-star

Habit Plants slender. Herbs, rhizomatous, not stoloniferous; caudex sometimes bearing bulbils.
Flowering stems

simple, 20–50 cm.

erect, leafy, 8–85 cm, stipitate-glandular.

Leaves

in basal rosette and cauline, basal digitately 3-lobed, segments again lobed, cauline (2–3), 3-foliolate or deeply lobed, reduced, similar to basal (except lobes longer);

stipules large, decurrent on petiole base;

petiole to 11 cm;

blade dark green or reddish green, orbiculate, (base hastate), surfaces densely hairy.

in basal rosette and cauline;

cauline leaves (1–)2(–10), usually alternate (opposite in L. cymbalaria), unlobed, lobed, or palmately compound, sometimes with bulbils in axils;

stipules present;

petiole stipitate-glandular;

blade rhombic or orbiculate to reniform, lobed, base cordate, hastate, rounded, ligulate, or cuneate, ultimate margins entire or toothed, apex obtuse or acute, surfaces often stipitate-glandular;

venation palmate.

Inflorescences

usually 1, (lax), nodding, 4–8-flowered racemes, simple.

compact or lax racemes, arising from axillary buds in basal rosette, 2–12(–25)-flowered, sometimes flowers solitary, (flowers sometimes replaced by bulbils), bracteate.

Pedicels

not exceeding length of hypanthium.

Flowers

deciduous if unfertilized, fragrant, horizontal;

hypanthium elongate-obconic, elongating in fruit, abruptly expanding, open at throat, (9–11 mm, length 3–4 times diam.);

sepals erect, triangular;

petals widely spreading, pink, obovate-rhombic, narrowly clawed, 3-lobed, 4–14 mm, ultimate margins entire;

ovary 1/2+ to ± entirely inferior;

styles included in fruit;

stigma papillae in narrow subapical band.

hypanthium partially adnate to proximal 1/2 to entire length of ovary, free from ovary 1–2 mm, green;

sepals 5, green tinged with red;

petals 5, white or pink; nectariferous tissue inconspicuous;

stamens 10, (inserted on hypanthium, inner whorl attached proximal to point of petal insertion);

filaments filiform; (anthers cordate);

pistil 3-carpellate;

ovary superior to 1/2 or almost completely inferior, 1-locular, carpels fully connate at ovaries;

placentation parietal;

styles 3;

stigmas 3.

Capsules

3-beaked.

Seeds

0.6–0.7 mm, smooth or wrinkled.

(50–200), usually dark brown, ovoid, smooth, wrinkled, or tuberculate.

x

= 7.

2n

= 28.

Lithophragma trifoliatum

Lithophragma

Phenology Flowering Mar–May.
Habitat Exposed runnels or near small streams, oak-coniferous woodland
Elevation 40-700 m (100-2300 ft)
Distribution
from FNA
CA
from USDA
w North America
[BONAP county map]
Discussion

Lithophragma trifoliatum is known from the western slope of the Cascade Range and Sierra Nevada in California. It is closely related to L. parviflorum and was considered a variety by P. E. Elvander (1993). The shape of the hypanthium, the fragrant flowers, the relatively large pink petals, and the relatively large seeds are distinctive. Lithophragma trifoliatum rarely produces seed; in cultivation, self-pollination was unsuccessful (R. L. Taylor 1965). It may represent a sterile derivative of L. parviflorum that now persists by vegetative reproduction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Tellima R. Brown [unranked] Lithophragma Nuttall, J. Acad. Nat. Sci. Philadelphia 7: 26. 1834 (as Lithofragma); Pleurendotria Rafinesque, name rejected

Species 10 (10 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves palmately compound, 3-foliolate; seeds spiny.
L. maximum
1. Leaves unlobed or lobed, or 3-foliolate; seeds smooth, wrinkled, tuberculate, or rough
→ 2
2. Stigma papillae in narrow subapical band; seeds smooth or wrinkled
→ 3
2. Stigma papillae covering apical surface; seeds tuberculate, tubercles in 3-19 rows, blunt or spinelike
→ 6
3. Leaf blades light green; cauline leaves 3-lobed, appearing pinnatifid; flowers persistent, slightly pendulous; racemes erect; petals 5-7-lobed; ovaries to 1/2 inferior.
L. tenellum
3. Leaf blades dark green or reddish green; cauline leaves 3-foliolate or 3-lobed; flowers deciduous if unfertilized, horizontal; racemes often lax, nodding; petals 3-lobed; ovaries 1/2+ inferior
→ 4
4. Hypanthia elongate-obconic, length 3-4 times diam.; petals pink; flowers fragrant.
L. trifoliatum
4. Hypanthia obconic to obconic-elongate, length 2 times diam.; petals usually white, sometimes pink; flowers not fragrant
→ 5
5. Cauline leaves 3-foliolate; hypanthia obconic-elongate; petals white or pink, often with prominent venation, 3-lobed, obovate-rhombic, widely spreading, ultimate margins entire; ovaries 1/2+ inferior.
L. parviflorum
5. Cauline leaves 3-lobed; hypanthia obconic; petals white with obscure venation, usually shallowly 3-lobed, ovate-elliptic, not widely spreading, ultimate margins sometimes with extra tooth, (lobes acute); ovaries ± completely inferior.
L. affine
6. Cauline leaves opposite (1 pair); flowering stems simple; petals unlobed, without serrations at base, ultimate margins entire.
L. cymbalaria
6. Cauline leaves alternate; flowering stems often branched; petals lobed or, if unlobed, margins sometimes with serrations at base
→ 7
7. Hypanthia broadly campanulate; flowers pendulous, petals partly included, ovate-elliptic, ultimate margins lacerate; flowers ± sessile.
L. campanulatum
7. Hypanthia narrowly campanulate; flowers horizontal, petals completely exserted, obovate-pandurate, ovate-elliptic, or ovate and palmately 5-lobed, ultimate margins entire; flowers long-pedicellate
→ 8
8. Hypanthia with truncate base; petals white, simple or 3-7-lobed; flowers not fragrant; racemes lax, nodding; cauline leaves 2-10; ovaries superior.
L. heterophyllum
8. Hypanthia with acute, obtuse, or hemispheric base; petals white or pink, unlobed, serrately 3-5-lobed, or 5-lobed; flowers sometimes fragrant; racemes erect; cauline leaves 2-4; ovaries to 1/2 inferior
→ 9
9. Plants usually robust; leaves usually unlobed or 3-5-lobed; petals white, unlobed or 3-5-lobed with serrations at base; inflorescences 2(-3)-branched, 3-5(-25)-flowered racemes, flowers rarely replaced with bulbils.
L. bolanderi
9. Plants usually fragile; leaves lobed (lobes rounded); petals usually pink, rarely white, palmately 5-lobed, base without serrations; inflorescences rarely branched (unless plant with bulbils), solitary flowers or 2-5(-7)-flowered racemes, often appearing corymbose; flowers sometimes replaced with bulbils.
L. glabrum
Source FNA vol. 8, p. 80. FNA vol. 8, p. 77. Author: Roy L. Taylor.
Parent taxa Saxifragaceae > Lithophragma Saxifragaceae
Sibling taxa
L. affine, L. bolanderi, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum
Subordinate taxa
L. affine, L. bolanderi, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
Synonyms L. parviflorum var. trifoliatum
Name authority Eastwood: Bull. Torrey Bot. Club 32: 200. 1905 (as trifoliata), (Nuttall) Torrey & A. Gray: Fl. N. Amer. 1: 583. 1840, name conserved ,
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