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mission star, mission woodland-star

Bolander's woodland-star, hillstar

Habit Plants slender. Plants usually robust.
Flowering stems

simple, 20–40 cm.

branched, (rarely bearing bulbils in place of flowers), 25–85 cm.

Leaves

in basal rosette and cauline, basal weakly 3-lobed, cauline (2), opposite, (1 pair), deeply 3-lobed, reduced, similar to basal;

stipules narrow, not decurrent on petiole, (margins fimbriate);

petiole to 9 cm;

blade dark green or reddish green, reniform, (base ligulate), surfaces sparsely hairy.

in basal rosette and cauline, basal unlobed or 3–5-lobed, never deeply lobed, cauline (2–3+), alternate, often 3–4-lobed, much reduced, similar to basal, (rarely with axillary bulbils);

stipules small, not decurrent on petiole, (margins fimbriate);

petiole usually to 25 cm;

blade green, orbiculate, surfaces hairy.

Inflorescences

solitary flowers or 1–3, (lax), nodding, 2–5(–8)-flowered racemes, simple, (10–40 cm).

1–3, erect, 3–5(–25)-flowered racemes, 2(–3)-branched, (10–85 cm).

Pedicels

1.5–2 times longer than hypanthium.

equaling or shorter than hypanthium, (flowers long-pedicellate).

Flowers

persistent, fragrant, vertical;

hypanthium turbinate, elongating in fruit, open at throat, (length 2 times diam.);

sepals erect, becoming wide-spreading after anthesis, triangular;

petals (exserted), widely spreading, white, ovate, narrowly clawed, unlobed, (bowl-shaped), 4–8 mm, ultimate margins entire;

ovary 1/2 inferior;

styles included in fruit;

stigma papillae apical.

persistent, fragrant, horizontal;

hypanthium narrowly campanulate with obtuse base, usually not elongating in fruit, throat open, (length 2 times diam.);

sepals erect in bud, widely spreading after anthesis, triangular;

petals (completely exserted), widely spreading, white, ovate-elliptic, narrowly clawed, usually unlobed with serrations at base, sometimes 3–5-lobed, 4–7 mm, ultimate margins entire;

ovary to 1/2 inferior;

styles slightly exerted in fruit;

stigma papillae apical.

Seeds

0.4 mm, tuberculate (tubercles in 3–19 rows, blunt or spinelike).

0.3–5 mm, tuberculate (tubercles in 3–19 rows, spinelike).

2n

= 14 + 1.

= 14, 28, 35, 42.

Lithophragma cymbalaria

Lithophragma bolanderi

Phenology Flowering Mar–Apr. Flowering May–Jun.
Habitat Shady oak-manzanita/arbutus woodland, mountainous regions, drainage courses Well-drained, open, grassy areas of oak or coniferous-oak woodland
Elevation 0-2000 m (0-6600 ft) 0-1500 m (0-4900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Lithophragma cymbalaria is the only species of the genus with a single pair of opposite cauline leaves. It shows a high degree of self-compatibility, with abundant seeds produced; it may produce abundant bulbils. It is found from Stanislaus County to Santa Barbara County including the northern Channel Islands.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Lithophragma bolanderi is known from California along the western slope of the Sierra Nevada from Shasta County to the San Gabriel Mountains and the north-central inner coastal mountains around San Francisco Bay north to Mendocino County.

Variable vegetative features, such as petal shape, size of seeds, and general growth habit, are the result of environmental factors and gene interchange among species. Lithophragma bolanderi is best distinguished from L. heterophyllum by the shape of the hypanthium base, position of styles and stigmas in fruit, length of seed tubercles, and general outline of the petals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 81. FNA vol. 8, p. 82.
Parent taxa Saxifragaceae > Lithophragma Saxifragaceae > Lithophragma
Sibling taxa
L. affine, L. bolanderi, L. campanulatum, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
L. affine, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
Synonyms Tellima cymbalaria L. heterophyllum var. bolanderi, L. heterophyllum var. scabrellum, L. scabrellum, L. scabrellum var. peirsonii, Tellima heterophylla var. bolanderi, Tellima scabrella
Name authority Torrey & A. Gray: Fl. N. Amer. 1: 585. 1840 , A. Gray: Proc. Amer. Acad. Arts 6: 535. 1865 ,
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