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Bolander's woodland-star, hillstar

slender fringecup, slender prairie star, slender woodland-star, woodland star

Habit Plants usually robust. Plants slender.
Flowering stems

branched, (rarely bearing bulbils in place of flowers), 25–85 cm.

simple, 15–30 cm.

Leaves

in basal rosette and cauline, basal unlobed or 3–5-lobed, never deeply lobed, cauline (2–3+), alternate, often 3–4-lobed, much reduced, similar to basal, (rarely with axillary bulbils);

stipules small, not decurrent on petiole, (margins fimbriate);

petiole usually to 25 cm;

blade green, orbiculate, surfaces hairy.

in basal rosette and cauline, basal unlobed, irregularly 3–5-lobed, or digitately lobed, sometimes almost pinnatifid, cauline (2), deeply 3-lobed, appearing pinnatifid, much reduced, more highly dissected than basal;

stipules broad, decurrent on petiole base, (margins fimbriate);

petiole to 8 cm;

blade light green, orbiculate, (base hastate), surfaces sparsely hairy.

Inflorescences

1–3, erect, 3–5(–25)-flowered racemes, 2(–3)-branched, (10–85 cm).

1–5, (compact), erect, 3–12-flowered racemes, simple, (10–12 cm).

Pedicels

equaling or shorter than hypanthium, (flowers long-pedicellate).

shorter than hypanthium.

Flowers

persistent, fragrant, horizontal;

hypanthium narrowly campanulate with obtuse base, usually not elongating in fruit, throat open, (length 2 times diam.);

sepals erect in bud, widely spreading after anthesis, triangular;

petals (completely exserted), widely spreading, white, ovate-elliptic, narrowly clawed, usually unlobed with serrations at base, sometimes 3–5-lobed, 4–7 mm, ultimate margins entire;

ovary to 1/2 inferior;

styles slightly exerted in fruit;

stigma papillae apical.

persistent, not fragrant, slightly pendulous;

hypanthium campanulate or hemispheric, becoming elongate-campanulate in fruit, open at throat;

sepals erect in bud, widely spreading after anthesis, triangular;

petals (exserted), widely spreading, pink, sometimes white, ovate, narrowly clawed, palmately 5–7-lobed, (sinuses extending 1/3–1/2 distance to base), 3–7 mm, ultimate margins entire;

ovary to 1/2 inferior;

styles exserted slightly in fruit;

stigma papillae in narrow subapical band.

Seeds

0.3–5 mm, tuberculate (tubercles in 3–19 rows, spinelike).

0.6–0.7 mm, smooth or wrinkled.

2n

= 14, 28, 35, 42.

= 14, 35.

Lithophragma bolanderi

Lithophragma tenellum

Phenology Flowering May–Jun. Flowering (Feb-)May–Jun(-Aug).
Habitat Well-drained, open, grassy areas of oak or coniferous-oak woodland Sagebrush desert in mountainous regions or high plateaus, dry, open forest, meadows, grasslands, and shrublands, sometimes in mesic subalpine habitats with Lithophragma parviflorum and L. glabrum
Elevation 0-1500 m (0-4900 ft) 1200-3000(-3400) m (3900-9800(-11200) ft)
Distribution
from FNA
CA
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from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; BC
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[BONAP county map]
Discussion

Lithophragma bolanderi is known from California along the western slope of the Sierra Nevada from Shasta County to the San Gabriel Mountains and the north-central inner coastal mountains around San Francisco Bay north to Mendocino County.

Variable vegetative features, such as petal shape, size of seeds, and general growth habit, are the result of environmental factors and gene interchange among species. Lithophragma bolanderi is best distinguished from L. heterophyllum by the shape of the hypanthium base, position of styles and stigmas in fruit, length of seed tubercles, and general outline of the petals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Lithophragma tenellum usually occurs on the eastern side of the Cascade Mountains and in the Rocky Mountains, Nevada, and Utah into western North America.

Taxonomy of Lithophragma tenellum is poorly understood because there are few collections from widely divergent geographical areas. The northwestern population (Washington, British Columbia) has been separated as a distinct species (L. thompsonii) based on the extent of the basal leaf lobation, which often shows considerable variation in all species. However, other populations in the Rocky Mountains, Nevada, and Utah have been observed with this lobation, as has Washington-British Columbia material having the more typical leaf form.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 82. FNA vol. 8, p. 79.
Parent taxa Saxifragaceae > Lithophragma Saxifragaceae > Lithophragma
Sibling taxa
L. affine, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
L. affine, L. bolanderi, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. trifoliatum
Synonyms L. heterophyllum var. bolanderi, L. heterophyllum var. scabrellum, L. scabrellum, L. scabrellum var. peirsonii, Tellima heterophylla var. bolanderi, Tellima scabrella L. australe, L. brevilobum, L. rupicola, L. tenellum var. thompsonii, L. thompsonii, Tellima tenella
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 535. 1865 , Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 584. 1840 (as tenella) ,
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