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fringecup, lithophragma, prairiestar, woodland-star

hillside woodland-star, woodland star

Habit Herbs, rhizomatous, not stoloniferous; caudex sometimes bearing bulbils. Plants slender.
Flowering stems

erect, leafy, 8–85 cm, stipitate-glandular.

simple, 20–40 cm.

Leaves

in basal rosette and cauline;

cauline leaves (1–)2(–10), usually alternate (opposite in L. cymbalaria), unlobed, lobed, or palmately compound, sometimes with bulbils in axils;

stipules present;

petiole stipitate-glandular;

blade rhombic or orbiculate to reniform, lobed, base cordate, hastate, rounded, ligulate, or cuneate, ultimate margins entire or toothed, apex obtuse or acute, surfaces often stipitate-glandular;

venation palmate.

in basal rosette and cauline, basal shallowly 2–5-lobed, cauline (2–10), 3-lobed with secondary teeth, usually much reduced, similar to basal, (bearing axillary bulbils, aerial bulbils may replace flowers in axils);

stipules small, not decurrent onto petiole, (margins fimbriate);

petiole to 8 cm;

blade dark green or reddish green, orbiculate, (margins toothed), surfaces hairy.

Inflorescences

compact or lax racemes, arising from axillary buds in basal rosette, 2–12(–25)-flowered, sometimes flowers solitary, (flowers sometimes replaced by bulbils), bracteate.

1–4(–6), (lax), nodding, 2–6(–12)-flowered racemes, often many-branched, (15–50 cm).

Pedicels

equaling or shorter than hypanthium, (flowers long-pedicellate).

Flowers

hypanthium partially adnate to proximal 1/2 to entire length of ovary, free from ovary 1–2 mm, green;

sepals 5, green tinged with red;

petals 5, white or pink; nectariferous tissue inconspicuous;

stamens 10, (inserted on hypanthium, inner whorl attached proximal to point of petal insertion);

filaments filiform; (anthers cordate);

pistil 3-carpellate;

ovary superior to 1/2 or almost completely inferior, 1-locular, carpels fully connate at ovaries;

placentation parietal;

styles 3;

stigmas 3.

persistent, not fragrant, horizontal;

hypanthium narrowly campanulate with truncate base, not elongating in fruit, throat open, (length 2 times diam.);

sepals erect in bud, becoming widely spreading after anthesis, triangular;

petals (completely exserted), widely spreading, (lax), white, obovate-pandurate, lamina spoon-shaped, narrowly clawed, unlobed or 3(–5)-lobed, 5–12 mm, ultimate margins entire;

ovary superior;

styles much exserted in fruit;

stigma papillae apical.

Capsules

3-beaked.

Seeds

(50–200), usually dark brown, ovoid, smooth, wrinkled, or tuberculate.

0.4–0.5 mm, tuberculate (tubercles in 3–19 rows, blunt).

x

= 7.

2n

= 14.

Lithophragma

Lithophragma heterophyllum

Phenology Flowering Jun.
Habitat Partly or fully shaded, well-drained slopes of oak or mixed coniferous-oak woodland
Elevation 0-1500 m (0-4900 ft)
Distribution
from USDA
w North America
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Tellima R. Brown [unranked] Lithophragma Nuttall, J. Acad. Nat. Sci. Philadelphia 7: 26. 1834 (as Lithofragma); Pleurendotria Rafinesque, name rejected

Species 10 (10 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Lithophragma heterophyllum is known in the coastal mountains of southern Oregon and in California from Humboldt to Los Angeles counties. Its variable morphology is associated with the amount of available moisture and site exposure.

Sympatric populations of Lithophragma heterophyllum and L. bolanderi occur in the south-central California Coast Ranges. The distinguishing characteristics for L. heterophyllum are hypanthium shape, petal outline, and exsertion of stigma and style in fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves palmately compound, 3-foliolate; seeds spiny.
L. maximum
1. Leaves unlobed or lobed, or 3-foliolate; seeds smooth, wrinkled, tuberculate, or rough
→ 2
2. Stigma papillae in narrow subapical band; seeds smooth or wrinkled
→ 3
2. Stigma papillae covering apical surface; seeds tuberculate, tubercles in 3-19 rows, blunt or spinelike
→ 6
3. Leaf blades light green; cauline leaves 3-lobed, appearing pinnatifid; flowers persistent, slightly pendulous; racemes erect; petals 5-7-lobed; ovaries to 1/2 inferior.
L. tenellum
3. Leaf blades dark green or reddish green; cauline leaves 3-foliolate or 3-lobed; flowers deciduous if unfertilized, horizontal; racemes often lax, nodding; petals 3-lobed; ovaries 1/2+ inferior
→ 4
4. Hypanthia elongate-obconic, length 3-4 times diam.; petals pink; flowers fragrant.
L. trifoliatum
4. Hypanthia obconic to obconic-elongate, length 2 times diam.; petals usually white, sometimes pink; flowers not fragrant
→ 5
5. Cauline leaves 3-foliolate; hypanthia obconic-elongate; petals white or pink, often with prominent venation, 3-lobed, obovate-rhombic, widely spreading, ultimate margins entire; ovaries 1/2+ inferior.
L. parviflorum
5. Cauline leaves 3-lobed; hypanthia obconic; petals white with obscure venation, usually shallowly 3-lobed, ovate-elliptic, not widely spreading, ultimate margins sometimes with extra tooth, (lobes acute); ovaries ± completely inferior.
L. affine
6. Cauline leaves opposite (1 pair); flowering stems simple; petals unlobed, without serrations at base, ultimate margins entire.
L. cymbalaria
6. Cauline leaves alternate; flowering stems often branched; petals lobed or, if unlobed, margins sometimes with serrations at base
→ 7
7. Hypanthia broadly campanulate; flowers pendulous, petals partly included, ovate-elliptic, ultimate margins lacerate; flowers ± sessile.
L. campanulatum
7. Hypanthia narrowly campanulate; flowers horizontal, petals completely exserted, obovate-pandurate, ovate-elliptic, or ovate and palmately 5-lobed, ultimate margins entire; flowers long-pedicellate
→ 8
8. Hypanthia with truncate base; petals white, simple or 3-7-lobed; flowers not fragrant; racemes lax, nodding; cauline leaves 2-10; ovaries superior.
L. heterophyllum
8. Hypanthia with acute, obtuse, or hemispheric base; petals white or pink, unlobed, serrately 3-5-lobed, or 5-lobed; flowers sometimes fragrant; racemes erect; cauline leaves 2-4; ovaries to 1/2 inferior
→ 9
9. Plants usually robust; leaves usually unlobed or 3-5-lobed; petals white, unlobed or 3-5-lobed with serrations at base; inflorescences 2(-3)-branched, 3-5(-25)-flowered racemes, flowers rarely replaced with bulbils.
L. bolanderi
9. Plants usually fragile; leaves lobed (lobes rounded); petals usually pink, rarely white, palmately 5-lobed, base without serrations; inflorescences rarely branched (unless plant with bulbils), solitary flowers or 2-5(-7)-flowered racemes, often appearing corymbose; flowers sometimes replaced with bulbils.
L. glabrum
Source FNA vol. 8, p. 77. Author: Roy L. Taylor. FNA vol. 8, p. 82.
Parent taxa Saxifragaceae Saxifragaceae > Lithophragma
Sibling taxa
L. affine, L. bolanderi, L. campanulatum, L. cymbalaria, L. glabrum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
Subordinate taxa
L. affine, L. bolanderi, L. campanulatum, L. cymbalaria, L. glabrum, L. heterophyllum, L. maximum, L. parviflorum, L. tenellum, L. trifoliatum
Synonyms Tellima heterophylla, L. trilobum
Name authority (Nuttall) Torrey & A. Gray: Fl. N. Amer. 1: 583. 1840, name conserved , (Hooker & Arnott) Hooker & Arnott: in J. Torrey and A. Gray, Fl. N. Amer. 1: 584. 1840 (as heterophylla) ,
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