Lithophragma |
Lithophragma heterophyllum |
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fringecup, lithophragma, prairiestar, woodland-star |
hillside woodland-star, woodland star |
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Habit | Herbs, rhizomatous, not stoloniferous; caudex sometimes bearing bulbils. | Plants slender. | ||||||||||||||||||||||||||||||||||||
Flowering stems | erect, leafy, 8–85 cm, stipitate-glandular. |
simple, 20–40 cm. |
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Leaves | in basal rosette and cauline; cauline leaves (1–)2(–10), usually alternate (opposite in L. cymbalaria), unlobed, lobed, or palmately compound, sometimes with bulbils in axils; stipules present; petiole stipitate-glandular; blade rhombic or orbiculate to reniform, lobed, base cordate, hastate, rounded, ligulate, or cuneate, ultimate margins entire or toothed, apex obtuse or acute, surfaces often stipitate-glandular; venation palmate. |
in basal rosette and cauline, basal shallowly 2–5-lobed, cauline (2–10), 3-lobed with secondary teeth, usually much reduced, similar to basal, (bearing axillary bulbils, aerial bulbils may replace flowers in axils); stipules small, not decurrent onto petiole, (margins fimbriate); petiole to 8 cm; blade dark green or reddish green, orbiculate, (margins toothed), surfaces hairy. |
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Inflorescences | compact or lax racemes, arising from axillary buds in basal rosette, 2–12(–25)-flowered, sometimes flowers solitary, (flowers sometimes replaced by bulbils), bracteate. |
1–4(–6), (lax), nodding, 2–6(–12)-flowered racemes, often many-branched, (15–50 cm). |
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Pedicels | equaling or shorter than hypanthium, (flowers long-pedicellate). |
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Flowers | hypanthium partially adnate to proximal 1/2 to entire length of ovary, free from ovary 1–2 mm, green; sepals 5, green tinged with red; petals 5, white or pink; nectariferous tissue inconspicuous; stamens 10, (inserted on hypanthium, inner whorl attached proximal to point of petal insertion); filaments filiform; (anthers cordate); pistil 3-carpellate; ovary superior to 1/2 or almost completely inferior, 1-locular, carpels fully connate at ovaries; placentation parietal; styles 3; stigmas 3. |
persistent, not fragrant, horizontal; hypanthium narrowly campanulate with truncate base, not elongating in fruit, throat open, (length 2 times diam.); sepals erect in bud, becoming widely spreading after anthesis, triangular; petals (completely exserted), widely spreading, (lax), white, obovate-pandurate, lamina spoon-shaped, narrowly clawed, unlobed or 3(–5)-lobed, 5–12 mm, ultimate margins entire; ovary superior; styles much exserted in fruit; stigma papillae apical. |
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Capsules | 3-beaked. |
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Seeds | (50–200), usually dark brown, ovoid, smooth, wrinkled, or tuberculate. |
0.4–0.5 mm, tuberculate (tubercles in 3–19 rows, blunt). |
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x | = 7. |
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2n | = 14. |
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Lithophragma |
Lithophragma heterophyllum |
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Phenology | Flowering Jun. | |||||||||||||||||||||||||||||||||||||
Habitat | Partly or fully shaded, well-drained slopes of oak or mixed coniferous-oak woodland | |||||||||||||||||||||||||||||||||||||
Elevation | 0-1500 m (0-4900 ft) | |||||||||||||||||||||||||||||||||||||
Distribution |
w North America |
CA; OR
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Discussion | Tellima R. Brown [unranked] Lithophragma Nuttall, J. Acad. Nat. Sci. Philadelphia 7: 26. 1834 (as Lithofragma); Pleurendotria Rafinesque, name rejected Species 10 (10 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lithophragma heterophyllum is known in the coastal mountains of southern Oregon and in California from Humboldt to Los Angeles counties. Its variable morphology is associated with the amount of available moisture and site exposure. Sympatric populations of Lithophragma heterophyllum and L. bolanderi occur in the south-central California Coast Ranges. The distinguishing characteristics for L. heterophyllum are hypanthium shape, petal outline, and exsertion of stigma and style in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 8, p. 77. | FNA vol. 8, p. 82. | ||||||||||||||||||||||||||||||||||||
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Synonyms | Tellima heterophylla, L. trilobum | |||||||||||||||||||||||||||||||||||||
Name authority | (Nuttall) Torrey & A. Gray: Fl. N. Amer. 1: 583. 1840, name conserved , | (Hooker & Arnott) Hooker & Arnott: in J. Torrey and A. Gray, Fl. N. Amer. 1: 584. 1840 (as heterophylla) , | ||||||||||||||||||||||||||||||||||||
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