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purple toad flax

butter-and-eggs, butter-and-eggs toadflax, common toadflax, greater butter-and-eggs, linaire vulgaire, yellow toadflax

Habit Perennials, from taproot, not reproducing vegetatively by stolons. Perennials, reproducing vegetatively by adventitious buds from roots.
Fertile stems

erect or suberect, to 71(–140) cm;

sterile stems to 18 cm.

erect or suberect, (25–)35–90(–120) cm;

sterile stems to 18 cm, usually absent.

Leaves

of fertile stems: blade oblanceolate to linear, usually flat, 5–45(–60) × 0.8–4(–8) mm, apex acute or subobtuse.

of fertile stems: blade linear to linear-lanceolate, 4–68 × 1–7(–15) mm, apex acute.

Racemes

1–117-flowered, dense;

bracts linear, 2–5(–5.5) × 0.3–1 mm.

11–31(–50)-flowered, usually dense, rachis glabrous or glandular-hairy, hairs 0.1–0.2 mm;

bracts linear to lanceolate, 4–20(–25) × 1–2 mm.

Pedicels

erect, 1–3 mm in flower, 2–4(–5) mm in fruit.

erect, 3–7.7 mm in flower, 3.5–8(–13) mm in fruit.

Styles

simple;

stigma entire.

simple;

stigma entire.

Corollas

violet to purple, with yellow or lilac palate, 9–13(–17) mm;

tube 1.5–2.5 mm wide, spurs curved, 5–7(–9) mm, subequal to rest of corolla, abaxial lip sinus (0.6–)0.8–1.5(–2) mm, adaxial lip sinus 1 mm.

white, pale yellow, or bright yellow, sometimes with orange palate, 27–32(–33) mm;

tube 5.5–9(–10) mm wide, spurs straight or curved, 11–15 mm, slightly shorter or subequal to rest of corolla, abaxial lip sinus 3–3.5 mm, adaxial lip sinus (2.2–)2.4–4(–4.5) mm.

Calyx

lobes linear to linear-lanceolate, 1.5–3 × 0.5–1 mm in flower, 2–3.5 × 0.7–1.2 mm in fruit, apex acute or subacute.

lobes lanceolate to ovate, 2.5–4.5 × 0.8–1.5 mm in flower, 2.5–5(–6) × 1–2(–2.5) mm in fruit, apex acute.

Capsules

subglobular, 2.7–4 × 2.5–3.7 mm, glabrous;

loculi equal.

oblong-globular or ovoid, 5–10(–11) × 3–6 mm, glabrous;

loculi equal.

Seeds

black or blackish brown, subtrigonous or ± tetrahedral, 0.8–1.2 × 0.6–1 mm, with longitudinal marginal ridges and anastomosed ridges or tubercles on faces;

wing absent.

gray, brown, or black, disciform, subglobular, with prominent encircling wing, 1.6–2.5(–3) × 1.5–2.3 mm, ± plane or biconvex;

disc gray, brown, or black, ± reniform, tuberculate;

wing ± entire, 0.4–0.7 mm wide, membranous, smooth.

2n

= 12 (Europe).

= 12.

Linaria purpurea

Linaria vulgaris

Phenology Flowering Jun–Oct. Flowering May–Oct.
Habitat Disturbed places, railroad rights-of-way, beach foreshores. Disturbed places, roadsides, railroad rights-of-way, rocky slopes, cultivated or abandoned fields, usually in moist or dry soils.
Elevation 0–1900 m. (0–6200 ft.) 0–2300(–3100) m. (0–7500(–10200) ft.)
Distribution
from FNA
CA; OR; WA; BC; s Europe (Italy) [Introduced in North America; introduced also in s South America (Argentina), n Europe, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Europe; c Asia [Introduced in North America; introduced also in temperate regions of s South America (Argentina), Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Linaria vulgaris was introduced into the northeastern United States (New England) before 1672 as an ornamental and medicinal plant (M. L. Fernald 1905b). In part from vigorous formation of adventitious shoots from the roots (T. S. Bakshi and R. T. Coupland 1960), L. vulgaris is now considered a noxious weed in nine states and four Canadian provinces (E. A. Lehnhoff 2008).

Hybrids between Linaria repens and L. vulgaris are known as L. ×sepium G. J. Allman; this hybrid presents an intermediate morphology between L. repens and L. vulgaris with regard to corolla and spur size (16–21 mm and 6–9.5 mm, respectively). Specimens examined suggest that this hybrid is relatively common in Newfoundland.

Hybrids between Linaria dalmatica subsp. dalmatica and L. vulgaris (L. ×hybrida Schur) are found in California (Lassen Creek, Modoc County), Colorado (Jefferson County), Idaho (Coeur d’Alene Lake, Kootenai County), and Montana (Beaverhead-Deerlodge National Forest) (S. M. Ward et al. 2009). In these localities, both potential parental species are sympatric or nearly sympatric. Ward et al. confirmed that hybridization occurs between populations of L. dalmatica and L. vulgaris, and that the hybrid progeny are viable and fertile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 31. FNA vol. 17, p. 28.
Parent taxa Plantaginaceae > Linaria Plantaginaceae > Linaria
Sibling taxa
L. bipartita, L. dalmatica, L. genistifolia, L. grandiflora, L. maroccana, L. repens, L. spartea, L. triornithophora, L. vulgaris
L. bipartita, L. dalmatica, L. genistifolia, L. grandiflora, L. maroccana, L. purpurea, L. repens, L. spartea, L. triornithophora
Synonyms Antirrhinum purpureum Antirrhinum linaria
Name authority (Linnaeus) Miller: Gard. Dict. ed. 8, Linaria no. 5. (1768) Miller: Gard. Dict. ed. 8, Linaria no. 1. (1768)
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