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purple toad flax

large-flower linaria, large-flower toadflax

Habit Perennials, from taproot, not reproducing vegetatively by stolons. Perennials, reproducing vegetatively by adventitious buds or stolons.
Fertile stems

erect or suberect, to 71(–140) cm;

sterile stems to 18 cm.

erect, (30–)60–74(–100) cm;

sterile stems to 13.5 cm.

Leaves

of fertile stems: blade oblanceolate to linear, usually flat, 5–45(–60) × 0.8–4(–8) mm, apex acute or subobtuse.

of fertile stems: blade ovate to lanceolate, flat, 12–48(–65) × 3–18(–30) mm, apex acute.

Racemes

1–117-flowered, dense;

bracts linear, 2–5(–5.5) × 0.3–1 mm.

1–36-flowered, dense in flower, lax in fruit;

bracts ovate to lanceolate, 7–14(–20) × 2–7 mm.

Pedicels

erect, 1–3 mm in flower, 2–4(–5) mm in fruit.

erect-patent to erect, (0.5–)2–6(–15) mm in flower, 4–15(–21) mm in fruit.

Styles

simple;

stigma entire.

simple;

stigma entire.

Corollas

violet to purple, with yellow or lilac palate, 9–13(–17) mm;

tube 1.5–2.5 mm wide, spurs curved, 5–7(–9) mm, subequal to rest of corolla, abaxial lip sinus (0.6–)0.8–1.5(–2) mm, adaxial lip sinus 1 mm.

pale to bright yellow, (40–)41–52 mm;

tube (6–)7.5–12 mm wide, spurs straight or curved, 19–22(–23) mm, slightly shorter or subequal to rest of corolla, abaxial lip sinus 2.5–3.5(–4) mm, adaxial lip sinus 4–6 mm.

Calyx

lobes linear to linear-lanceolate, 1.5–3 × 0.5–1 mm in flower, 2–3.5 × 0.7–1.2 mm in fruit, apex acute or subacute.

lobes ovate, sometimes ovate-lanceolate, 7–14 × 3.5–7 mm in flower, 7–14 × 3.5–9 mm in fruit, apex acute.

Capsules

subglobular, 2.7–4 × 2.5–3.7 mm, glabrous;

loculi equal.

globular, 6–8(–11) × 6–10.7 mm, glabrous;

loculi equal.

Seeds

black or blackish brown, subtrigonous or ± tetrahedral, 0.8–1.2 × 0.6–1 mm, with longitudinal marginal ridges and anastomosed ridges or tubercles on faces;

wing absent.

black, subtrigonous or tetrahedral, 0.9–1.5 mm, with longitudinal acute marginal ridges and anastomosed acute ridges on faces;

wing absent.

2n

= 12 (Europe).

Linaria purpurea

Linaria grandiflora

Phenology Flowering Jun–Oct. Flowering Jul–Sep.
Habitat Disturbed places, railroad rights-of-way, beach foreshores. Roadsides, disturbed places.
Elevation 0–1900 m. (0–6200 ft.) 600–2100 m. (2000–6900 ft.)
Distribution
from FNA
CA; OR; WA; BC; s Europe (Italy) [Introduced in North America; introduced also in s South America (Argentina), n Europe, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; WA; se Europe; sw Asia [Introduced in North America]
Discussion

Linaria grandiflora is closely related to L. dalmatica and sometimes has been included within L. dalmatica. One specimen collected in Washington (Kittitas County, west of Cle Elum, 18 June 1962, C. L. Hitchcock & C. V. Muhlick 22329, NY) exhibits discordant features (for example, pedicels 15–21 mm). According to D. A. Sutton (1988), L. grandiflora bears pedicels 0.5–4 mm, although he indicated that plants called L. pancicii Janka (which were included in the synonymy of L. grandiflora) have long pedicels. For the rest of the characteristics (corolla size, adaxial lip sinus length, calyx lobe size and shape), that specimen is fairly assimilable in L. grandiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 31. FNA vol. 17, p. 30.
Parent taxa Plantaginaceae > Linaria Plantaginaceae > Linaria
Sibling taxa
L. bipartita, L. dalmatica, L. genistifolia, L. grandiflora, L. maroccana, L. repens, L. spartea, L. triornithophora, L. vulgaris
L. bipartita, L. dalmatica, L. genistifolia, L. maroccana, L. purpurea, L. repens, L. spartea, L. triornithophora, L. vulgaris
Synonyms Antirrhinum purpureum
Name authority (Linnaeus) Miller: Gard. Dict. ed. 8, Linaria no. 5. (1768) Desfontaines: Ann. Mus. Natl. Hist. Nat. 11: 51, plate 2. (1808)
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