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ambulia, marshweed

Habit Herbs, perennial [annual], paludal or aquatic, emergent.
Stems

erect [prostrate, creeping], hairy or glabrous.

Leaves

cauline, opposite or whorled, dimorphic;

petiole present or absent;

blade not fleshy, not leathery, margins of submerged leaves pinnatifid, of aerial leaves entire, serrate, or pinnatifid.

Inflorescences

terminal or axillary, racemes or flowers solitary, rarely paired [or spikes];

bracts present or absent.

Pedicels

present or absent;

bracteoles present or absent.

Flowers

bisexual, chasmogamous or cleistogamous;

sepals 5, basally connate, calyx bilaterally symmetric, tubular, lobes triangular or lanceolate [ovate];

corolla blue, blue-purple, or white, bilaterally symmetric, bilabiate, tubular or funnelform, tube base not gibbous or spurred, lobes 4 or 5, abaxial 3, adaxial 1 or 2;

stamens 4, basally adnate to corolla, didynamous, filaments glabrous or hairy;

staminode 0;

ovary 2-locular, placentation axile;

stigma capitate, 2-lobed.

Fruits

capsules, dehiscence septicidal.

Seeds

20–200, brown to dark brown, conic to cylindric, wings absent.

× = 17.

Limnophila

Distribution
from USDA
Asia; Africa; Pacific Islands; Australia [Introduced in North America; introduced also in South America]
[BONAP county map]
Discussion

Species 40 (2 in the flora).

The two species recognized here could be difficult to distinguish throughout their ranges; intermediate plants, which are believed to be hybrids, have been found.

Limnophila is closely related to Hydrotriche Zuccarini, a small genus of aquatic plants native to Madagascar. Together they form a group sister to Gratiola in Gratioleae (D. Estes and R. L. Small 2008).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pedicels 3–8 mm in flower, 5–12 mm in fruit; bracteoles 1–3.5 mm; corolla tubes 4–5(–8) mm; cleistogamous flowers absent.
L. indica
1. Pedicels 0–2 mm in flower, 0–4 mm in fruit; bracteoles 0–1.5 mm; corolla tubes 8–10 mm; cleistogamous flowers present, submerged.
L. sessiliflora
Source FNA vol. 17, p. 271. Author: Kerry A. Barringer.
Parent taxa Plantaginaceae
Subordinate taxa
L. indica, L. sessiliflora
Name authority R. Brown: Prodr., 442. (1810) — name conserved
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