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woolly meadowfoam

Photo is of parent taxon

Bellinger's meadowfoam

Habit Plants 3–25 cm; herbage glabrous or sparsely to densely hairy.
Herbage

glabrous.

Stems

erect to ascending or decumbent.

erect to ascending.

Leaves

1–8 cm;

leaflets 5–11, blade linear to ovate-elliptic, margins entire, irregularly toothed, or lobed.

Flowers

urn-, cup-, or bell-shaped;

sepals accrescent or not, ovate, obovate, lanceolate, oblong-lanceolate, or lanceolate-ovate, 4–10 mm;

petals white, obovate, oblong, or obovate-cuneate, 4.5–10 mm, 1.6–1.8 times as long as wide, 0.5–1.1 times longer than sepals, apex retuse, obtuse, erose, truncate, or emarginate;

filaments 2–7 mm;

anthers (yellow) 0.4–1.5 mm;

style 1.5–4 mm.

urn-shaped;

sepals accrescent, narrowly ovate, 5.5–8 mm, abaxially and adaxially glabrous or sparsely hairy;

petals oblong, 5.5–7.5 mm, apex retuse, (without marginal hairs basally);

filaments 2–3.5 mm;

anthers ± 0.5 mm, usually dehiscing introrsely;

style 2.3–2.5 mm.

Nutlets

dark brown or gray, 3–4.5 mm, tuberculate, tubercles straw-colored, platelike, conic, or awl-shaped.

1 or 2 (3), tubercles platelike.

Limnanthes floccosa

Limnanthes floccosa subsp. bellingeriana

Phenology Flowering Apr–May.
Habitat Vernal pool edges in shallow soil of rocky meadows
Elevation 300-1200 m [1000-3900 ft]
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

D. Southworth and J. Seevers (1997) compared pollen of subspp. bellingeriana and floccosa. Pollen of Limnanthes floccosa has a striated zone perpendicular to the apertural band; in subsp. bellingeriana, the striations are criss-crossed and in subsp. floccosa the striations are parallel. Southworth and Seevers reported that the two subspecies could be easily distinguished in the field in mixed populations based on the density of hairs on the sepals. In subsp. floccosa, the hairs are so dense that they prevent the sepals from fully opening; in subsp. bellingeriana, the sepal hairs are relatively short (50 µm) and sparse. Based on the absence of intermediates in these characters, Southworth and Seevers recommended elevating subsp. bellingeriana to species level. Allozyme studies (C. I. McNeill and S. K. Jain 1983) group subspp. bellingeriana and floccosa together, and subspp. californica, grandiflora, and pumila together. Also, in ITS analysis (M. S. Plotkin 1998), subsp. bellingeriana is nested within the floccosa clade. Crosses between subspp. bellingeriana and floccosa resulted in viable hybrids (R. Ornduff 1971). Subspecies bellingeriana is found in shaded, damp edges of meadows; sympatric subsp. floccosa occurs on the upper, dry, exposed edges of meadows (M. T. K. Arroyo 1973). On balance, I prefer to retain this taxon as a subspecies within L. floccosa. Subspecies bellingeriana is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowers bell- or urn-shaped (petals without marginal hairs basally); anthers usually dehiscing introrsely
→ 2
1. Flowers cup-shaped (petals with marginal hairs basally); anthers usually dehiscing extrorsely (introrsely in subsp. pumila)
→ 3
2. Sepals abaxially and adaxially densely villous; nutlet tubercles awl-shaped.
subsp. floccosa
2. Sepals abaxially and adaxially glabrous or sparsely hairy; nutlet tubercles platelike.
subsp. bellingeriana
3. Herbage densely hairy; sepals abaxially densely hairy.
subsp. californica
3. Herbage glabrous or sparsely hairy; sepals abaxially glabrous or sparsely hairy
→ 4
4. Sepals not accrescent, 7.5-8 mm, adaxially glabrous.
subsp. pumila
4. Sepals accrescent, 8.5-9 mm, adaxially densely hairy.
subsp. grandiflora
Source FNA vol. 7, p. 182. FNA vol. 7, p. 182.
Parent taxa Limnanthaceae > Limnanthes > sect. Inflexae Limnanthaceae > Limnanthes > sect. Inflexae > Limnanthes floccosa
Sibling taxa
L. alba, L. bakeri, L. douglasii, L. macounii, L. montana, L. vinculans
L. floccosa subsp. californica, L. floccosa subsp. floccosa, L. floccosa subsp. grandiflora, L. floccosa subsp. pumila
Subordinate taxa
L. floccosa subsp. bellingeriana, L. floccosa subsp. californica, L. floccosa subsp. floccosa, L. floccosa subsp. grandiflora, L. floccosa subsp. pumila
Synonyms L. bellingeriana
Name authority Howell: Fl. N.W. Amer., 108. (1897) (M. Peck) C. T. Mason: Univ. Calif. Publ. Bot. 25: 501. (1952)
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