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Cascade lily, Mount Hood lily, Shasta lily, Washington lily

Kelley's lily

Bulbs

subrhizomatous to ± ovoid, 2.7–10 × 4.4–13.4 cm, 0.2–0.9(–1.4) times taller than long;

scales unsegmented or notched with 2(–3) poorly defined segments, sometimes 2-segmented, longest 3.3–11.9 cm;

stem roots absent.

rhizomatous, unbranched, continuously scaly, 1.4–3.1 × 3.8–8.2 cm, 0.3–0.6 times taller than long;

scales 2–3-segmented, a few unsegmented, longest 0.8–2.7 cm;

stem roots absent.

Stems

to 2 m, often glaucous.

to 2.2 m. Buds rounded in cross section.

Buds

rounded in cross section.

Leaves

in 1–8 whorls or partial whorls, 3–16 leaves per whorl, horizontal and drooping at tips to ascending, occasionally nearly clasping stem, 3.7–12.3 × 0.9–4.7 cm, 2–6.5 times longer than wide;

blade oblanceolate, sometimes obovate, rarely elliptic, margins undulate or not, apex acute, often widely so;

veins and margins ± smooth abaxially.

scattered or usually in 1–4 whorls or partial whorls, 3–10 leaves per whorl, horizontal and drooping at tips, 7.8–15.8 × 1.1–5.3 cm, 3.2–10.8 times longer than wide;

blade usually elliptic, occasionally weakly lanceolate or oblanceolate, margins not undulate, apex narrowly acute;

veins and margins ± smooth abaxially.

Inflorescences

racemose, 1–33-flowered.

racemose, 1–15(–25)-flowered.

Flowers

opening before dusk, ± horizontal, slightly bilaterally symmetric, strongly fragrant;

perianth ± funnelform;

sepals and petals recurved 2/3–3/4 along length from base and scarcely overlapping to form tube, lower usually less recurved than upper and forming landing platform, white, often aging deep pink or lavender, often with short yellowish stripe extending from basal median nectaries, often with fine magenta spots concentrated basally, not distinctly clawed;

sepals sometimes purplish abaxially, not ridged abaxially, (6.1–)6.7–11.3 × 0.9–1.7 cm;

petals noticeably wider than sepals, oblanceolate and often very wide distally, (6.1–)6.6–11.2 × 1.1–2.4 cm, apex widely acute, obtuse, or rounded;

stamens barely exserted;

filaments barely spreading, diverging 2°–8° from axis;

anthers off white or cream, becoming pale pink or yellow, 0.8–1.5 cm;

pollen yellow or cream;

pistil 7.5–10.4 cm;

ovary 1.7–3.4 cm;

style pale green;

pedicel 4.8–15 cm.

pendent, mildly fragrant;

perianth Turk’s-cap-shaped;

sepals and petals reflexed 2/5 along length from base, yellow or yellow-orange, usually with copious maroon spots, not distinctly clawed;

sepals not ridged abaxially, 4.3–5.7 × 0.9–1.5 cm;

petals 4.2–5.7 × 0.9–1.5 cm;

stamens moderately exserted;

filaments moderately spreading, diverging 9°–19° from axis;

anthers magenta or dull red, 0.3–0.6 cm;

pollen pale orange, rust, orange, or tan-orange;

pistil 2.6–3.4 cm;

ovary 0.8–1.4 cm;

style green;

pedicel 8.5–20.7 cm.

Capsules

often with 6 longitudinal ridges, 2.7–5.8 × 1.6–2.9 cm, 1.3–2.3 times longer than wide.

1.5–2.9 × 1–1.9 cm, 1.1–2 times longer than wide.

Seeds

123–231.

96–169.

2n

= 24.

Lilium washingtonianum

Lilium kelleyanum

Phenology Flowering summer (Jul–Aug).
Habitat Hillside seeps, wet thickets, along streams in coniferous forests
Elevation 2200–2900 m (7200–9500 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Lilium kelleyanum has in common with L. parvum a short, relatively wide fruit and small flowers, and divides with that species the montane and subalpine Sierra Nevada on both sides of the divide from Lake Tahoe south. The two species are parapatrically distributed with an intervening zone of introgression; intergrades occur, among other places, in the June Lake region about ten miles south of Mono Lake in Mono County and in adjacent Madera County. The two species appear to be sister taxa (M. W. Skinner 1988), which argues against subsuming L. kelleyanum as a subspecies of L. pardalinum, as has been suggested by several students of the group. Despite the close affinity of L. kelleyanum to L. parvum, taxonomic custom within the genus dictates that they be retained as separate species since floral features are well differentiated.

Lilium kelleyanum is pollinated by swallowtail butterflies (J. S. Davis 1956).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Sepals and petals aging light pink or remaining white; bulbs sometimes (36%) without notched or segmented scales; sepals 8–11.3 cm; Sierra Nevada and s Cascades of California.
subsp. washingtonianum
1. Sepals and petals aging deep pink or lavender; bulbs usually (88%) with some notched or segmented scales; sepals (6.1–)6.7–9.5 cm; mountains of n California and Oregon.
subsp. purpurascens
Source FNA vol. 26, p. 182. FNA vol. 26, p. 187.
Parent taxa Liliaceae > Lilium Liliaceae > Lilium
Sibling taxa
L. bolanderi, L. canadense, L. catesbaei, L. columbianum, L. grayi, L. humboldtii, L. iridollae, L. kelleyanum, L. kelloggii, L. lancifolium, L. maritimum, L. michauxii, L. michiganense, L. occidentale, L. pardalinum, L. parryi, L. parvum, L. philadelphicum, L. pyrophilum, L. rubescens, L. superbum
L. bolanderi, L. canadense, L. catesbaei, L. columbianum, L. grayi, L. humboldtii, L. iridollae, L. kelloggii, L. lancifolium, L. maritimum, L. michauxii, L. michiganense, L. occidentale, L. pardalinum, L. parryi, L. parvum, L. philadelphicum, L. pyrophilum, L. rubescens, L. superbum, L. washingtonianum
Subordinate taxa
L. washingtonianum subsp. purpurascens, L. washingtonianum subsp. washingtonianum
Synonyms L. fresnense, L. inyoense, L. nevadense var. fresnense, L. nevadense var. monense, L. pardalinum var. parviflorum
Name authority Kellogg: Hesperian (San Francisco) 3: 340. (1859) Lemmon: Sierra Club Bull. 4: 300. (1903)
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