Lilium rubescens |
Lilium humboldtii |
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chaparral lily, redwood lily |
Humboldt lily, Humboldt's lily |
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Bulbs | ± ovoid, 3.9–9.4 × 2.5–7.1 cm, 1.1–2.6 times taller than long; scales unsegmented, longest 4–8.9 cm; stem roots absent. |
often somewhat purplish, variable, subrhizomatous to ± ovoid, 3.3–11.7 × 2.4–14.4 cm, 0.4–2.6(–3) times taller than long; scales unsegmented or notched, longest 3.5–11.2 cm; stem roots present or absent. |
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Stems | to 2 m, often glaucous. |
to 3.1 m. Buds rounded in cross section. |
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Buds | rounded in cross section. |
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Leaves | in 3–5(–more) whorls or partial whorls, 3–15 leaves per whorl, usually ± ascending, sometimes horizontal and drooping at tips, 3.2–12.3 × 1–2.8 cm, 2–6.4 times longer than wide; blade oblanceolate, sometimes obovate, rarely elliptic, margins usually undulate, apex acute, often widely so, or obtuse; veins and margins ± smooth abaxially. |
in 2–8 whorls or partial whorls, 3–16(–28) leaves per whorl, usually ascending, 4.6–14.5 × 0.8–3.6 cm, 2.9–7.2 times longer than wide; blade usually ± oblanceolate, though often weakly so, rarely elliptic or lanceolate, margins usually undulate, apex acute; veins and margins ± smooth abaxially. |
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Inflorescences | racemose, often with flowers in loosely defined whorls, 1–40-flowered. |
racemose, 1–33(–40)-flowered. |
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Flowers | ascending to erect, fragrant; perianth funnelform; sepals and petals recurved 1/2–2/3 along length from base, white, aging pink-purple, with fine magenta spots concentrated proximally or less often generally distributed, often reddish or purplish abaxially, not distinctly clawed; sepals not ridged abaxially, 4.3–6.6 × 0.6–1.4 cm; petals noticeably wider than sepals, oblanceolate and often very wide proximally, 4.2–6.4 × 0.9–1.9 cm, apex widely acute, obtuse, or rounded; stamens included; filaments barely spreading, diverging 3°–12° from axis; anthers pale yellow, 0.4–0.8 cm; pollen yellow; pistil 2.7–3.8 cm; ovary 1.2–1.8 cm; style pale green; pedicel 1.4–9.5 cm. |
pendent, not fragrant; perianth Turk’s-cap-shaped; sepals and petals reflexed 1/5 along length from base, orange or yellow, with prominent red or magenta spots, not distinctly clawed; sepals not ridged abaxially, 5.2–9.8 × 1.4–2.4 cm; petals 5.6–9.6 × 1.5–2.9 cm; stamens strongly exserted; filaments parallel along most of length, then very widely spreading, diverging 17°–31° from axis; anthers purple, 1.1–1.9 cm; pollen rust, tan, or peach, becoming yellow or tan-yellow; pistil 4.6–7.1 cm; ovary 1.2–2.6 cm; style green, often pale; pedicel 7.8–21.2 cm. |
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Capsules | usually with 6 longitudinal ridges, 2–3.7 × 1.7–2.7 cm, 1.1–1.7 times longer than wide. |
longitudinally keeled, 2.5–5.4 × 1.8–3.3 cm, 1.1–2.3 times longer than wide. |
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Seeds | 67–189. |
not counted. |
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2n | = 24. |
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Lilium rubescens |
Lilium humboldtii |
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Phenology | Flowering summer (late May–early Aug). | |||||
Habitat | Dry soils in chaparral, gaps in redwood [Sequoia sempervirens (D. Don) Endlicher] or mixed evergreen forests | |||||
Elevation | 0–1500 m (0–4900 ft) | |||||
Distribution |
CA
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CA
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Discussion | Lilium rubescens occurs from Santa Cruz County north through the Coast Ranges to Del Norte County, and is declining in the southern part of its range due to urbanization and over-collecting. This is the only species in the genus in which the chromosome complement is known to consist of one pair of long metacentric chromosomes and eleven pairs of acrocentric chromosomes; all others possess two long metacentric pairs and ten shorter acrocentric pairs (M. W. Skinner 1988). Various bee species visit the flowers during the day to collect nectar and pollen and may be the most significant pollinators. Pale swallowtails (Papilio eurymedon Lucas, family Papilionidae) also visit this lily. The Karok used Lilium rubescens decoratively in bouquets (D. E. Moerman 1986). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Lilium humboldtii is declining throughout its range due to habitat destruction, primarily for housing. Forming large scattered colonies at foothill elevations under ponderosa pines or in oak canyons and chaparral, these massive plants with towering inflorescences and large flowers are quite striking. With Lilium pardalinum and L. parryi, they were used in the early part of the century to produce the Bellingham hybrid lilies, development of which continued with the Bellmaid hybrids. Though not as popular as various Asiatic hybrids, these are still in use. A. M. Kellogg was aware as early as 1859 that the tall, orange-flowered, dry-land lily from the foothills of the Sierra Nevada was distinctive, and he presented this opinion to the California Academy of Sciences about that time (A. M. Kellogg 1872). By the time he had published a description attached to the name Lilium bloomerianum, P. E. S. Duchartre had described this same plant from material provided by B. Roezl and grown by M. Leichtlin, and named it after the German explorer and botanist Alexander von Humboldt. Carl Purdy and others then misapplied Kellogg’s name in various combinations to the southern California expression recognized here as Lilium humboldtii subsp. ocellatum. The Humboldt lily is pollinated primarily by large butterflies, especially western tiger swallowtails (Papilio rutulus Lucas, family Papilionidae) and pale swallowtails (P. eurymedon Lucas), and to a lesser extent by the monarch butterfly [Danaus plexippus (Linnaeus), family Nymphalidae]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 183. | FNA vol. 26, p. 181. | ||||
Parent taxa | Liliaceae > Lilium | Liliaceae > Lilium | ||||
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Synonyms | L. washingtonianum var. purpureum | |||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 14: 256. (1879) | Roezl & Leichtlin ex Duchartre: J. Soc. Centr. Hort. France, sér. 2, 5: 43. (1871) | ||||
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