Lilium humboldtii |
Lilium parvum |
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Humboldt lily, Humboldt's lily |
alpine lily, Sierra tiger lily |
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Bulbs | often somewhat purplish, variable, subrhizomatous to ± ovoid, 3.3–11.7 × 2.4–14.4 cm, 0.4–2.6(–3) times taller than long; scales unsegmented or notched, longest 3.5–11.2 cm; stem roots present or absent. |
rhizomatous, unbranched, continuously scaly, 1.4–3.5 × 3.3–9.2 cm, 0.3–0.5 times taller than long; scales (1–)2–3(–4)-segmented, longest 1.1–3.4 cm; stem roots absent. |
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Stems | to 3.1 m. Buds rounded in cross section. |
to 1.7 m. Buds rounded in cross section. |
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Leaves | in 2–8 whorls or partial whorls, 3–16(–28) leaves per whorl, usually ascending, 4.6–14.5 × 0.8–3.6 cm, 2.9–7.2 times longer than wide; blade usually ± oblanceolate, though often weakly so, rarely elliptic or lanceolate, margins usually undulate, apex acute; veins and margins ± smooth abaxially. |
in 2–5 whorls or partial whorls, 3–13 leaves per whorl, ± horizontal and drooping at tips or ascending in sun, 4–15.1 × 0.5–4.4 cm, 2.3–10.8 times longer than wide; blade ± elliptic, margins not undulate, apex acute, often narrowly so; veins and margins ± smooth abaxially. |
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Inflorescences | racemose, 1–33(–40)-flowered. |
racemose, 1–26(–41)-flowered. |
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Flowers | pendent, not fragrant; perianth Turk’s-cap-shaped; sepals and petals reflexed 1/5 along length from base, orange or yellow, with prominent red or magenta spots, not distinctly clawed; sepals not ridged abaxially, 5.2–9.8 × 1.4–2.4 cm; petals 5.6–9.6 × 1.5–2.9 cm; stamens strongly exserted; filaments parallel along most of length, then very widely spreading, diverging 17°–31° from axis; anthers purple, 1.1–1.9 cm; pollen rust, tan, or peach, becoming yellow or tan-yellow; pistil 4.6–7.1 cm; ovary 1.2–2.6 cm; style green, often pale; pedicel 7.8–21.2 cm. |
± horizontal to ascending, sometimes slightly bilaterally symmetric, not fragrant; perianth ± funnelform; sepals and petals somewhat recurved 3/5–2/3 along length from base, not strongly reflexed, lower often less recurved than upper and forming landing platform, orange or yellowish proximally, darker (light orange, orange, red-orange, red, or rarely ± pinkish) on distal 2/5, occasionally uniformly light orange or rarely yellow, spotted dark purple-brown or maroon, often pale abaxially, not distinctly clawed; sepals not ridged abaxially, noticeably wider than petals, 3.2–4.2 × 0.8–1.3 cm; petals 3.2–4.2 × 0.6–1.2 cm; stamens moderately exserted; filaments moderately spreading, diverging 8°–17° from axis; anthers pale yellow, orangish, or magenta, 0.3–0.8 cm; pollen yellowish to red-orange; pistil 2.3–3.7 cm; ovary 0.8–1.4 cm; style pale green; pedicel 3.5–15.5 cm. |
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Capsules | longitudinally keeled, 2.5–5.4 × 1.8–3.3 cm, 1.1–2.3 times longer than wide. |
1.6–2.7 × 1.2–1.8 cm, 1.2–1.9 times longer than wide. |
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Seeds | not counted. |
90–132. |
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2n | = 24. |
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Lilium humboldtii |
Lilium parvum |
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Phenology | Flowering summer (mid Jun–Aug). | |||||
Habitat | Wet meadows, willow (Salix spp.) thickets, and streams in coniferous forests | |||||
Elevation | 1400–2900 m (4600–9500 ft) | |||||
Distribution |
CA
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CA; NV
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Discussion | Subspecies 2 (2 in the flora). Lilium humboldtii is declining throughout its range due to habitat destruction, primarily for housing. Forming large scattered colonies at foothill elevations under ponderosa pines or in oak canyons and chaparral, these massive plants with towering inflorescences and large flowers are quite striking. With Lilium pardalinum and L. parryi, they were used in the early part of the century to produce the Bellingham hybrid lilies, development of which continued with the Bellmaid hybrids. Though not as popular as various Asiatic hybrids, these are still in use. A. M. Kellogg was aware as early as 1859 that the tall, orange-flowered, dry-land lily from the foothills of the Sierra Nevada was distinctive, and he presented this opinion to the California Academy of Sciences about that time (A. M. Kellogg 1872). By the time he had published a description attached to the name Lilium bloomerianum, P. E. S. Duchartre had described this same plant from material provided by B. Roezl and grown by M. Leichtlin, and named it after the German explorer and botanist Alexander von Humboldt. Carl Purdy and others then misapplied Kellogg’s name in various combinations to the southern California expression recognized here as Lilium humboldtii subsp. ocellatum. The Humboldt lily is pollinated primarily by large butterflies, especially western tiger swallowtails (Papilio rutulus Lucas, family Papilionidae) and pale swallowtails (P. eurymedon Lucas), and to a lesser extent by the monarch butterfly [Danaus plexippus (Linnaeus), family Nymphalidae]. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
At the lower elevations within its range, Lilium parvum tends to have clear yellow-orange flowers—a form that has been recognized as L. parvum var. crocatum—but perianth parts are redder apically at higher elevations in the Sierra Nevada. The pink-flowered form that is apparently localized at lower elevations in El Dorado County, California, is referred to informally as the ditch lily (O. H. Ballantyne 1983). Lilium parvum hybridizes sporadically with L. pardalinum subsp. pardalinum at middle elevations (ca. 1200–1500 m) in the Sierra Nevada, resulting in variable swarms of plants with flowers that are intermediate in orientation, size, and perianth recurvature. Hybrids with Lilium kelleyanum are discussed under that species. Lilium parvum is a pollination generalist visited by several species of hummingbirds (family Trochilidae), western tiger swallowtails (Papilio rutulus Lucas, family Papilionidae), pale swallowtails (P. eurymedon Lucas), and various bees (mostly family Apidae). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 181. | FNA vol. 26, p. 188. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | L. canadense var. parvum, L. parvum var. crocatum | |||||
Name authority | Roezl & Leichtlin ex Duchartre: J. Soc. Centr. Hort. France, sér. 2, 5: 43. (1871) | Kellogg: Hesperian (San Francisco) 8: 163. (1862) | ||||
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