Leymus triticoides |
Poaceae subfam. pooideae |
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beardless lyme grass, beardless wildrye, creeping wildrye |
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Habit | Plants not cespitose, strongly rhizomatous. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 45-125 cm tall, 1.8-3 mm thick, solitary or few together. |
usually hollow, sometimes solid. |
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Leaves | exceeded by the spikes, often basally concentrated; sheaths glabrous or hairy, hairs 0.5-1 mm; auricles to 1 mm; ligules 0.2-1.3 mm, truncate, erose; blades 10-35 cm long, 3.5-10 mm wide, flat to involute, usually stiffly ascending, adaxial surfaces usually scabrous, often also sparsely hairy, hairs to 0.8 mm, most abundant proximally, veins 11-27, closely spaced, subequal, prominently ribbed. |
distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikes | 5-20 cm long, 5-15 mm wide, with 2 spikelets at midspike, sometimes 1 or 3 at other nodes; internodes 5-11.5 mm, usually mostly smooth and glabrous, sometimes strigillose distally, edges ciliate, cilia to 0.4 mm. |
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Spikelets | 10-22 mm, with 3-7 florets. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | 5-16 mm long, 0.5-1.2 mm wide, bases not overlapping, glabrous and smooth proximally, scabrous distally, tapering from below midlength to the subulate apices, stiff, keeled, the central portion thicker than the margins, 1(3)-veined, veins inconspicuous at midlength; calluses usually glabrous, occasionally with a few hairs, hairs about 0.1 mm; lemmas 5-12 mm, usually glabrous, occasionally sparsely hairy, hairs to 0.3 mm, apices acute, usually awned, awns to 3 mm; anthers 3-6 mm, dehiscent. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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2n | = 28. |
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Leymus triticoides |
Poaceae subfam. pooideae |
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Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; TX; UT; WA; WY; HI; BC
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Discussion | Leymus triticoides grows in dry to moist, often saline meadows. Its range extends from southern British Columbia to Montana, south to California, Arizona, and New Mexico, but its populations are widely scattered. It is not known from Mexico. There is considerable variation within the species, but no pattern of variation suggesting the existence of infraspecific taxa is known. It is very similar to L. multicaulis, strains of which were initially released as L. triticoides by the U.S. Department of Agriculture. The most consistent differences between them appear to be in the venation of the leaf blades and the vestiture of the calluses. Leymus triticoides is also very similar to L. simplex, differing from it in the number of spikelets at the midspike nodes. Leymus triticoides hybridizes with other species of Leymus; hybrids with L. mollis are called L. xvancouverensis (see p. 358), those with L. condensatus are called L. xmultiflorus (see p. 362). Hybrids with L. cinereus are known, but have not been formally named. Plants identified as Elymus arenicolus Scribn. & J.G. Sm. are here included in L. flavescens, but may represent hybrids between L. triticoides and L. flavescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 360. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Triticeae > Leymus | Poaceae | ||||||||||||||||||||||||||||||||||||
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Synonyms | Elymus triticoides var. pubescens, Elymus triticoides | |||||||||||||||||||||||||||||||||||||
Name authority | (Buckley) Pilg. | Benth. | ||||||||||||||||||||||||||||||||||||
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