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American dunegrass, American lyme grass, dune wildrye, dunegrass, elyme des sables d'amerique, sea lyme-grass, seigle de mer

Habit Plants not cespitose, strongly rhizomatous, occasionally slightly glaucous. Plants cespitose, sometimes weakly rhizomatous.
Culms

12-170 cm tall, 3-6 mm thick, usually densely pubescent below the spikes for 10-40+ mm.

35-140 cm tall, 1.5-3 mm thick, several together.

Leaves

exceeded by the spikes;

auricles to 0.7 mm;

ligules 0.2-2.5 mm;

blades 10-94 cm long, 3-15 mm wide, adaxial surfaces scabridulous to scabrous, 20-40-veined, veins subequal, prominently ribbed, closely spaced.

exceeded by the spikes;

auricles to 1 mm;

ligules 0.1-1 mm, truncate;

blades 1-5 mm wide, flat to strongly involute, adaxial surfaces glabrous or sparsely to -9 prominently ribbed, subequal long, 4-11 mm wide, nodes below midspike with 1-2(3) spikelets, distal nodes with 1 spikelet;

internodes 3.5-9 mm, surfaces glabrous, edges scabrous or strigillose.

Spikes

5-34 cm long, 10-20 mm wide, with 3-33 nodes, usually with 2 spikelets per node;

internodes 4.5-9.5 mm, surfaces and edges similar, hairs on the surfaces 0.1-0.5 mm, on the edges to 0.7 mm.

Spikelets

15-34 mm, with 3-6 florets.

9-21 mm, pedicellate, pedicels to 1 mm, with 3-6 florets.

Glumes

9-34 mm long, 1.5-4 mm wide, lanceolate, tapering from midlength or above, flat or rounded on the back, flexible, usually strigillose to pilose or villous, rarely almost glabrous, the central portion scarcely thicker than the margins, 3(5)-veined at midlength, apices acute;

lemmas 11-20 mm, densely hairy, hairs 0.5-1 mm, soft, apices acute, unawned;

anthers 4-9 mm, dehiscent.

unequal to subequal, to 12.5 mm long, 0.5-3.2 mm wide, subulate, stiff, keeled, the central portion thicker than the margins, tapering from below midlength, 0-1(3)-veined, veins inconspicuous at midlength;

lower glumes 0-12 mm;

upper glumes 3.5-12.5 mm;

lemmas 7-12.5 mm, usually glabrous, sometimes sparsely strigillose, unawned or awned, awns to 2.5 mm;

anthers 2.5-7.5 mm, dehiscent.

Leymus mollis

Leymus salina

Distribution
from FNA
AK; CA; IL; MA; ME; MI; NH; OR; PA; WA; WI; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
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Discussion

Leymus mollis is native to Asia and North America. It is treated here as having two very similar subspecies that have somewhat different ranges. The subspecies are sometimes treated as separate species, but they may be little more than environmentally induced variants. Both subspecies grow primarily on coastal beaches, close to the high tide line, and along some inland waterways, particularly in the arctic. Reports of L. ajanensis (V.N. Vassil.) Tzvelev from North America are based on specimens of L. mollis (D. Murray, University of Alaska, pers. comm. 2006).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The three subspecies of Leymus salina differ in their pubescence and geographic distribution, with subsp. salina being the most common of the three. The specific epithet comes from the locality of the type collection: Salina Pass, Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikes 12-34 cm long, with 12-33 nodes; basal blades 5-15 mm wide; culms 50-170 cm tall
subsp. mollis
1. Spikes 5-13(16) cm long, with 3-14 nodes; basal blades 3-8 mm wide; culms 12-70 cm tall
subsp. villosissimus
1. Basal sheaths and blades conspicuously hairy on the abaxial surfaces
subsp. salmonis
1. Basal sheaths glabrous; blades usually glabrous on the abaxial surfaces.
→ 2
2. Blades strongly involute, usually densely hairy just above the ligules
subsp. salina
2. Blades flat or almost flat, not densely hairy above the ligules
subsp. mojavensis
Source FNA vol. 24, p. 356. FNA vol. 24, p. 364.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Leymus Poaceae > subfam. Pooideae > tribe Triticeae > Leymus
Sibling taxa
L. ambiguus, L. angustus, L. arenarius, L. californicus, L. cinereus, L. condensatus, L. flavescens, L. innovatus, L. multicaulis, L. pacificus, L. racemosus, L. salina, L. simplex, L. triticoides, L. ×multiflorus, L. ×vancouverensis
L. ambiguus, L. angustus, L. arenarius, L. californicus, L. cinereus, L. condensatus, L. flavescens, L. innovatus, L. mollis, L. multicaulis, L. pacificus, L. racemosus, L. simplex, L. triticoides, L. ×multiflorus, L. ×vancouverensis
Subordinate taxa
L. mollis subsp. mollis, L. mollis subsp. villosissimus
L. salina subsp. mojavensis, L. salina subsp. salina, L. salina subsp. salmonis
Synonyms Elymus mollis, Elymus arenarius var. villosus, Elymus arenarius var. mollis, Elymus arenarius subsp. mollis Elymus salinus, Elymus ambiguus var. salina
Name authority (Trin.) Pilg. (M.E. Jones) Á. Löve
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