Cistanthe tweedyi(synonym of Lewisiopsis tweedyi) |
Cistanthe |
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Tweedy's bitterroot, Tweedy's lewisia, Tweedy's pussypaws |
pussypaws |
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Habit | Plants perennial, taprooted; caudex short-branching. | Herbs, annual or perennial, succulent, glabrous, with caudices, not rhizomatous or stoloniferous. | ||||||||||||||||||||||||||||||||||||||||
Roots | fleshy and/or fibrous, or taproots. |
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Stems | 1 or more from each rosette, 10–20 cm, bracts scattered proximally, ovate-lanceolate, scarious. |
decumbent to erect, simple or branched; nodes glabrous. |
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Leaves | basal, in loose rosettes; blade elliptic-lanceolate to ovate or oblanceolate, narrowing to broad petiole, 6–15 cm. |
basal and/or cauline, glabrous, not articulate at base, somewhat to markedly clasping, attachment points linear; basal leaves in rosettes; cauline leaves alternate (usually absent in C. umbellata and C. tweedyi). |
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Inflorescences | racemose, 1–5(–8)-flowered. |
axillary, umbellate, paniculate, racemose, or cymose, sometimes scorpioid and/or secund, loose to dense, 1–many-flowered, leafy and/or bracteate; bracts (1–)2 subtending each flower, markedly unequal. |
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Flowers | pedicellate; sepals broadly ovate, 9–10 mm, scarious; petals 7–9(–12), salmon-pink to yellowish, rarely white, 25–40 mm; stamens 10–23, anther yellow; style present; stigmas 3; pedicel 20–60 mm. |
pedicellate to subsessile; sepals persistent, distinct, not keeled or angled, often unequal, herbaceous to scarious, glabrous; petals 2–9(–12), distinct; stamens 1–23; ovules 1–many; style present or absent; stigmas 2 or 3. |
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Capsules | ovoid, 7–10 mm, dehiscence circumscissile near base; valves 3–4, splitting from base toward apex. |
2–3-valved, dehiscence loculicidal from apex, valves not deciduous, (except C. tweedyi, which has circumscissile deshiscence and 3–4 valves splitting from base), margins not reflexed after dehiscence, margins not markedly involute. |
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Seeds | 12–35, dark brownish red, suborbicular to reniform, 2 mm, not shiny, warty, strophiolate. |
1–40, dark brownish red, dull gray, or black, usually elliptic to orbiculate, sometimes orbiculate-reniform, dull or shiny, smooth or sculptured, sometimes hairy, strophiolate in C. maritima and C. tweedyi. |
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x | = 22, 23. |
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2n | = 46, 92. |
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Cistanthe tweedyi |
Cistanthe |
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Phenology | Flowering May–Jul. | |||||||||||||||||||||||||||||||||||||||||
Habitat | Well-drained granitic talus slopes and ledges, often in ponderosa pine duff | |||||||||||||||||||||||||||||||||||||||||
Elevation | 600-2200 m (2000-7200 ft) | |||||||||||||||||||||||||||||||||||||||||
Distribution |
WA; BC
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South America; North America (including Mexico) |
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 35 (11 in the flora). Publications by R. C. Carolin (1987) and M. A. Hershkovitz (1991, 1991b) have required the addition to the North American flora of the genus Cistanthe, which originally was erected to accommodate some Chilean species that were segregated from Calandrinia. Recent investigations indicate that a considerably broader range of species belongs in the genus, including two North American species formerly placed in Calandrinia, all the species formerly included in Calyptridium, and one species previously classified in Lewisia. While the current recognition of Cistanthe originally rested on the cladistic work of Carolin and the studies of leaf morphology by Hershkovitz, who documented nine traits of leaf morphology distinguishing Cistanthe, more recent molecular studies by M. A. Hershkovitz and E. A. Zimmer (2000) have provided general support for it. The inclusion of C. tweedyi appears to be somewhat equivocal and it might best be treated as a distinct genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 464. | FNA vol. 4, p. 460. | ||||||||||||||||||||||||||||||||||||||||
Parent taxa | Portulacaceae > Cistanthe | Portulacaceae | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | Calandrinia tweedyi, Lewisia tweedyi, Lewisiopsis tweedyi | Calyptridium, Lewisiopsis, Spraguea | ||||||||||||||||||||||||||||||||||||||||
Name authority | (A. Gray) Hershkovitz: Phytologia 68: 268. (1990) | Spach: Hist. Nat. Vég. 5: 229. (1836) | ||||||||||||||||||||||||||||||||||||||||
Web links |