Lewisia |
Lewisia nevadensis |
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bitterroot, lewisia |
Nevada bitterroot, Nevada lewisia, Sierra lewisia |
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Habit | Herbs, perennial, with enlarged caudices. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Roots | taproots, gradually ramified distally or fusiform to napiform, rarely globose and cormlike, fleshy. |
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Taproots | napiform to shortly fusiform. |
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Stems | deciduous, prostrate to erect, scapelike, simple or branched; nodes glabrous. |
suberect, becoming horizontal or deflexed after anthesis, base subterranean, 5–12 cm. |
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Leaves | basal or basal and cauline; basal leaves evergreen or ephemeral, in basal rosettes or tufts (except in L. triphylla); cauline leaves, if present, alternate, opposite, or whorled, ± sessile or gradually or abruptly narrowed basally into broad, clasping petiole; blade fleshy, margins entire, toothed, or crisped, often hyaline near base. |
basal leaves withering at or soon after anthesis, gradually narrowed to broad petiole, blade narrowly linear to linear-oblanceolate, flattened, 4–15 cm, margins entire, apex obtuse to subacute; cauline leaves absent. |
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Inflorescences | racemose, paniculate, or subumbellate cymes, pedunculate, or with flowers borne singly; bracts persistent, 2(–9) at each flowering node, subequal, margins entire, toothed, or glandular-toothed, herbaceous or scarious. |
usually with flowers borne singly, rarely 2–3-flowered in racemose cymes; bracts 2, opposite, linear-lanceolate, 6–18 mm, margins entire, apex acute. |
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Flowers | pedicellate or sessile, persistent or disarticulate in fruit; sepals persistent, 2(–9), equal or subequal when paired, margins entire, toothed, or glandular-toothed, herbaceous or scarious; petals (4–)5–10(–19), twisting about and falling away with capsules after anthesis; stamens 1–50, distinct to scarcely connate and/or adnate to petals basally; ovules 1–50; style branched; stigmas 2–8. |
pedicellate, not disarticulate in fruit, 0.5–2 cm diam.; sepals 2, broadly ovate, 5–13 mm, herbaceous at anthesis, margins entire or with few shallow, nonglandular teeth, apex acute to subacute; petals 5–10, white or rarely pinkish, elliptic to oblanceolate, 10–15(–20) mm; stamens 6–15; stigmas 3–6; pedicel 10–40 mm. |
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Capsules | dehiscence circumscissile near base, splitting from base toward apex. |
5–10 mm. |
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Seeds | 1–50, brown or black, smooth or minutely sculpted, estrophiolate. |
20–50, 1.3 mm, shiny, muricate. |
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x | = 10, 11, 12, 13, 14, 15. |
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2n | = 56. |
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Lewisia |
Lewisia nevadensis |
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Phenology | Flowering late spring–late summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Wet grassy slopes and meadows near springs | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1300-3200 m [4300-10500 ft] | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
w North America (including Mexico) |
AZ; CA; CO; ID; NM; NV; OR; UT; WA
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Discussion | Species 16 (16 in the flora). The number and circumscription of species in Lewisia are not widely agreed upon because of morphological variability and intergradation. Natural hybridization is inferred for some plants. In cultivation, the species reputedly freely intercross; most hybrid offspring are sterile. Most species are cultivated by alpine/succulent plant enthusiasts. In the wild, the plants usually occur in small, isolated populations in rocky or gravelly places, especially in canyons or alpine flats. The key to species here has been modified from B. Mathew (1989b). Hybrids, cultivars, and plants in cultivation may not key satisfactorily. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lewisia nevadensis represents one extreme of the L. pygmaea complex (see discussion under 13. L. pygmaea). Questionable geographic occurrences reflect plants that have one or more features otherwise suggestive of L. pygmaea (e.g., more elongate roots, truncate and/or toothed sepals, and colored petals); such intermediates also occur in the range of “typical” L. nevadensis (relatively robust plants with napiform roots, solitary flowers, acute sepals with entire margins, and white petals). Uncertainty respecting the affinity of specimens prevails in those from Arizona, Colorado, Idaho, and Wyoming. There are no supporting specimens from Wyoming. B. L. Davidson (2000) noted that Lewisia nevadensis is a garden weed in Colorado; it is not clear whether or not these plants are escapes from cultivation. The floral symmetry of Lewisia nevadensis may be somewhat elliptical, the two outer sepals and the remaining petals imbricate and opposite the sepals, giving the flowers a pinched appearance, a feature also reported for L. oppositifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 476. | FNA vol. 4. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Erocallis, Oreobroma | Calandrinia nevadensis, Claytonia grayana, L. bernardina, L. pygmaea var. nevadensis, Oreobroma nevadense | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Pursh: Fl. Amer. Sept. 2: 360. (1814) | (A. Gray) B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1: 268. (1897) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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