Leucolepis acanthoneura |
Leucolepis |
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leucolepis umbrella moss |
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Habit | Plants 3–8(–10) cm, in tufts or mats. | |
Stem(s) | leaves on young shoots and subtending branches green to pale green, those below branches reddish brown proximally and hyaline distally, slightly contorted and rarely somewhat crisped when dry, appressed to stem proximally and slightly spreading distally, ± flat when moist, (2–)2.5–3(–4) mm; margins hyaline or reddish brown, teeth large, more pronounced distally, often at right angles to margin; apex long-acuminate; medial laminal cells elongate, 20–50 µm; marginal cells linear and short-linear, in 1 or 2 rows. |
reddish brown to black, erect, branched distally, dendroid; rhizoids brown, macronemata matted proximally, occasionally along stem into branched portion of stem, micronemata absent. |
Leaves | dimorphic, pale to dark green or reddish brown to hyaline, slightly contorted or crisped when dry, appressed or spreading, flat or keeled when moist, ovate-lanceolate, (1.3–)2–3(–4) mm; base decurrent; margins plane, green, hyaline, or reddish brown, 1-stratose, usually toothed to near base, teeth single, sharp; apex acute or long-acuminate, cuspidate or not; costa subpercurrent or percurrent, distal abaxial surface strongly toothed; medial laminal cells elongate, short-elongate, short-rhombic, or ± isodiametric, 17–50 µm, sometimes in longitudinal or diagonal rows, not or weakly collenchymatous, walls not pitted; marginal cells differentiated, linear, short-linear, or rhomboidal, in 1 or 2 rows. |
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Branch leaves | green to dark green, moderately crisped when dry, spreading, keeled when moist, 1.3–2(–2.5) mm; margins green, teeth small, more pronounced distally, not at right angles to margin; apex acute, cuspidate; medial laminal cells short-elongate, short-rhombic, or ± isodiametric, 17–27 µm; marginal cells short-linear and rhomboidal, in 1 (or 2) rows. |
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Seta | single or rarely double (or 3), reddish brown, 4–5 cm, somewhat flexuose. |
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Sexual condition | dioicous. |
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Capsule | with annulus differentiated in 3 rows of cells; exostome papillose; endostome papillose, basal membrane porous, cilia 2 or 3. |
pendent, yellow-brown or brown, cylindric, 6–8 mm; operculum hemispheric; exostome yellow-brown; endostome yellow-brown. |
Spores | 28–30 µm. |
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Specialized | asexual reproduction absent. |
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Leucolepis acanthoneura |
Leucolepis |
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Phenology | Capsules mature spring–summer. | |
Habitat | Soil, boulders, rotten logs, tree trunks, shaded habitats, often alongside trails in lowlands and forests | |
Elevation | low to moderate elevations (0-1000 m) (low to moderate elevations (0-3300 ft)) | |
Distribution |
AK; CA; ID; OR; WA; BC |
w North America |
Discussion | Plants of Leucolepis acanthoneura have the appearance of small palm trees due to their dendroid growth form and recurved branches. The specific epithet alludes to spiny teeth on the abaxial surface of the costa. Climacium dendroides, another dendroid moss in the west, is distinguished by rhizomelike stems, erect capsules, and broad, cordate stem leaves with rounded apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 1. Leucolepis is distinguished by a dendroid growth form and strongly differentiated stem leaves. Morphological (T. J. Koponen 1968) and cytological (R. J. Lowry 1948; W. C. Steere et al. 1954) research supports the split of Leucolepis from Mnium. The chromosome number of Leucolepis is n = 5, whereas Mnium is n = 6, 7, and 12. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 222. | FNA vol. 28, p. 222. |
Parent taxa | Mniaceae > Leucolepis | Mniaceae |
Subordinate taxa | ||
Synonyms | Hypnum acanthoneuron, L. menziesii, Mnium menziesii | |
Name authority | (Schwagrichen) Lindberg: Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 81. (1868) | Lindberg: Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 80. (1868) |
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