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green spangletop, green sprangletop

sprangletop

Habit Plants perennial. Plants annual or perennial; cespitose.
Culms

(10) 30-110 cm, round or basally compressed, tillering from the basal nodes, not branching from the aerial nodes, mostly glabrous, sometimes pilose basally;

internodes solid.

(3)10-250(300) cm, usually ascending to erect, often geniculate at the lower nodes, occasionally prostrate and rooting at the lower nodes, often branching at the aerial nodes;

nodes usually glabrous;

internodes usually hollow.

Sheaths

sometimes with a pilose collar;

ligules 1-2 mm, truncate, erose;

blades (2)8-35 cm long, 2-8 mm wide, glabrous, strigose, or pilose.

Leaves

usually primarily cauline, occasionally in basal rosettes;

sheaths open;

ligules 0.2-10(15) mm, obtuse to attenuate, usually membranous, sometimes ciliate;

blades flat, involute when dry, usually ascending to erect, apices attenuate.

Panicles

8-20 cm, with 2-15 subdigitate or racemose branches;

secondary panicles often hidden in the lowest leaf sheaths;

branches 2-19 cm, ascending to spreading at maturity.

Spikelets

4-12 mm, light brown to dark olive green, with 4-13 florets, often widely diverging at anthesis.

rounded to slightly keeled, distant to tightly imbricate, not conspicuously pubescent, with (2)3-12(20) bisexual florets;

rachillas rarely prolonged.

Glumes

narrowly triangular to ovate, acute;

lower glumes 2.3-4.8 mm;

upper glumes 3.3-6 mm;

lemmas 3.5-5 mm, membranous, ovate to obovate, lateral veins glabrous or sericeous, hairs often restricted to the basal portion, sometimes also sericeous on the midvein and between the veins, apices obtuse to truncate, usually emarginate, unawned, sometimes mucronate;

paleas ciliate on the margins;

anthers 3, 0.3-1.6 mm.

usually unequal, sometimes subequal, exceeded by the florets, membranous, rounded to weakly keeled, 1-veined, veins scabrous, apices unawned (rarely mucronate);

lower glumes 0.5-4.9 mm;

upper glumes 0.9-6 mm;

florets usually bisexual;

calluses distinct or poorly developed, glabrous or pubescent;

lemmas membranous, usually pubescent at least over the lower portion of the veins, 3(5)-veined, apices entire or minutely bilobed, unawned, mucronate, or awned;

paleas usually subequal to the lemmas, membranous or hyaline;

anthers 1-3, 0.1-2.7 mm.

Caryopses

1.5-2.3 mm long, 0.9-1 mm wide, strongly dorsally compressed.

obovate to elliptic, falling free of the lemmas and paleas.

Primary

inflorescences terminal, panicles of 2-150 non-disarticulating, spikelike branches, usually exceeding the leaves;

branches 1-22 cm, digitate, subdigitate, or racemose on the rachises, 1-sided, usually spikelet-bearing throughout their length, spikelets in 2 rows, axes terminating in a functional spikelet, lower branches occasionally with secondary branching;

secondary panicles sometimes present, axillary to and concealed by the lower sheaths, their florets not disarticulating;

disarticulation in the primary panicles beneath the florets.

x

= 10.

2n

= 40, 60, 80.

Leptochloa dubia

Leptochloa

Distribution
from FNA
AZ; CA; CO; FL; KS; MD; MO; MS; NC; NM; OK; SC; TX
[WildflowerSearch map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; BC; ON; QC; Virgin Islands
[BONAP county map]
Discussion

Leptochloa dubia grows from the southwestern United States and Florida through Mexico to Argentina, often in well-drained, sandy or rocky soils. It provides fair to good forage, but is seldom abundant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Leptochloa is a pantropical, warm-temperate genus of 32 species. Eight of the ten species in this treatment are native to the Flora region. Of the other two, L. chloridiformis was introduced over 60 years ago but has not become established. Leptochloa chinensis is not yet known from the region; it is included here because of its potential threat as an invasive weed. Cladistic studies (Snow 1997) do not support recognition of the segregate genus Diplachne.

Leptochloa tends to grow in somewhat basic soils. Many of the species, particularly the annual species, are poor ecological competitors and grow in relatively open, seasonally inundated soils, such as are found along rivers. In disturbed areas, they are associated with roadside ditches, the margins of reservoirs, and mesic agricultural lands. A few species, primarily perennial, grow on well-drained soils. The vegetative vigor of all species is greatly influenced by soil moisture availability.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches digitate or subdigitate; plants perennial.
→ 2
2. Lemma apices obtuse to truncate and often emarginate; lemmas membranous; plants often with secondary panicles concealed in the lower leaf sheaths
L. dubia
2. Lemma apices usually acute; lemmas chartaceous; plants without secondary panicles concealed in the lower leaf sheaths.
→ 3
3. Panicle branches always digitate, 7-17 cm long; lemmas mucronate but not awned; in the Flora region, known only from a few old collections in Cameron County, Texas
L. chloridiformis
3. Panicle branches subdigitate, 1.5-18 cm long; lemmas unawned, mucronate, or awned; native in much of the southeastern United States, including parts of Texas
L. virgata
1. Panicle branches racemose; plants annual or perennial.
→ 4
4. Ligules 2-8 mm long, attenuate, becoming lacerate at maturity
L. fusca
4. Ligules 0.3-5.4 mm long, truncate to obtuse, erose, ciliate, or lacerate.
→ 5
5. Sheaths sparsely to densly hairy, the hairs papillose-based
L. panicea
5. Sheaths lacking hairs or with hairs that are not papillose-based.
→ 6
6. Panicles with 25-150 branches.
→ 7
7. Lemmas 2.4-3 mm long; anthers 0.6-0.8 mm long; spikelets 4-5 mm long
L. panicoides
7. Lemmas 1.2-2.4 mm long; anthers 0.2-0.6 mm long; spikelets 2.5-4.5 mm long.
→ 8
8. Leaf blades 8-16 mm wide; lemmas 2.1-2.4 mm long
L. scabra
8. Leaf blades 4-8 mm wide; lemmas 1-2 mm long.
→ 9
9. Lower glumes 0.7-0.8 mm long; anthers 0.2-0.4 mm long; plants native to the southern United States
L. nealleyi
9. Lower glumes 1.1-1.7 mm long; anthers 0.4-0.6 mm long; plants aggressive weeds, currently not known from the Flora region
L. chinensis
6. Panicles with 2-25 branches.
→ 10
10. Plants perennial.
→ 11
11. Lemmas 4-5 mm long, membranous, their apices broadly acute, obtuse, or truncate, unawned; panicles with 2-15 branches; caryopses 1.9-2.3 mm long, strongly dorsally compressed; secondary panicles often present in, and concealed by, the lower leaf sheaths
L. dubia
11. Lemmas 1.5-3.6 mm long, chartaceous, their apices usually acute, rarely obtuse, unawned, mucronate, or awned, the awns to 11 mm long; panicles with 9-25 branches; caryopses 1.3-1.8 mm long, somewhat laterally compressed; secondary panicles not present in the lower leaf sheaths
L. virgata
10. Plants annual.
→ 12
12. Panicles 2-17 cm long, with 5-23 branches; anthers 0.4-0.5 mm long; caryopses 0.4-0.5 mm wide
L. viscida
12. Panicles 20-35 cm long, with 20-90 branches; anthers 0.6-0.8 mm long; caryopses about 0.7 mm wide
L. panicoides
Source FNA vol. 25, p. 54. FNA vol. 25, p. 51. Author: Neil Snow;.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Leptochloa Poaceae > subfam. Chloridoideae > tribe Cynodonteae
Sibling taxa
L. chinensis, L. chloridiformis, L. fusca, L. nealleyi, L. panicea, L. panicoides, L. scabra, L. virgata, L. viscida
Subordinate taxa
L. chinensis, L. chloridiformis, L. dubia, L. fusca, L. nealleyi, L. panicea, L. panicoides, L. scabra, L. virgata, L. viscida
Name authority (Kunth) Nees P. Beauv.
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