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mesa pepperwort

hairypod pepperweed, shaggyfruit pepperweed

Habit Annuals, biennials, or perennials; (base woody); glabrous or pubescent. Annuals; hirsute or hispid, (trichomes cylindrical).
Stems

simple from base, erect, branched distally, (3.5–)4.5–16(–18) dm.

usually few to several, rarely simple from base, erect to ascending or (outer ones) decumbent, branched distally, (0.15–)0.6–3(–3.8) dm.

Basal leaves

(soon deciduous), not rosulate;

petiole (1–)2–5.5(–7.5) cm;

blade pinnatifid, (2–)3–6.8(–9) cm × 10–30 mm, margins (of lobes) dentate to serrate.

(later withered);

not rosulate;

petiole (0.4–)1–3.5(–5) cm;

blade spatulate to oblanceolate, lyrate-pinnatifid, pinnatisect, or 2-pinnatifid, (0.7–)1.5–4.5(–7.5) cm × (9–)12–20(–30) mm, margins rarely dentate, (lobes) entire or dentate.

Cauline leaves

shortly petiolate or sessile;

blade narrowly lanceolate or oblanceolate to linear, 3–7 cm × (2.5–)4–10mm (smaller distally), base attenuate to cuneate, not auriculate, margins usually entire, rarely dentate.

subsessile or petioles 0.8–2.2 cm, blade lanceolate to oblanceolate, (0.7–)1.2–3.3(–5) cm × (2–)4–12 mm, base cuneate, not auriculate, margins subentire to dentate.

Racemes

elongated in fruit;

rachis puberulent, trichomes straight or curved.

often considerably elongated in fruit;

rachis hirsute or hispid, trichomes straight, cylindrical.

Flowers

sepals suborbicular to oblong, 0.8–1.5 × 0.7–1.2 mm;

petals white, suborbicular, 2.2–3.5(4) × 1.5–2.5 mm, claw 0.7–1.5 mm;

stamens 6;

filaments 1.5–2.5 mm, (glabrous);

anthers 0.3–0.4 mm.

sepals oblong, 1–1.3(–1.5) × 0.5–0.8mm;

petals (sometimes absent), white, oblanceolate to linear, (0.3–)0.6–1.5(–2) × (0.1–)0.2–0.5 mm, claw absent;

stamens 2, median;

filaments 1–1.4 mm;

anthers 0.2–0.3 mm.

Fruiting pedicels

divaricate-ascending to horizontal, usually slightly recurved or somewhat sigmoid, rarely straight, (not winged), (3–)3.5–7.5(–8) × 0.2–0.3 mm, puberulent adaxially.

divaricate-ascending to horizontal, straight or slightly curved, (often strongly flattened), (1.8–)2–4(–4.6) × 0.2–0.7 mm (to 0.3 mm thick), hirsute to hispid throughout or adaxially.

Fruits

broadly ovate, 2–3.5(–4) × 1.8–2.6(–3) mm, apically winged, apical notch 0.1–0.2mm deep;

valves thin, smooth, not veined, glabrous;

style (0.2–)0.3–0.6(–0.7) mm, exserted beyond apical notch.

ovate to ovate-orbicular, 2.8–4(–4.6) × 2.4–3.6(–4) mm, (base broadly cuneate to rounded), apically winged, apical notch (0.2–)0.3–0.6(–0.7) mm deep;

valves thin, smooth, not veined, hirsute to hispid (on surface or margin);

style obsolete or to 0.1 mm, included in apical notch.

Seeds

(dark brown), ovate, 1.4–1.8 × 0.8–1.1 mm.

ovate, 1.4–2.2 × 0.9–1.4 mm.

Lepidium eastwoodiae

Lepidium lasiocarpum

Phenology Flowering Jul–Sep.
Habitat Pinyon-juniper, sagebrush, or mixed desert shrub communities
Elevation 900-2200 m (3000-7200 ft)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; NV; TX; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

C. L. Hitchcock (1936) and R. C. Rollins (1993) reduced Lepidium eastwoodiae to a variety of L. montanum and L. alyssoides, respectively. However, the differences in morphology and flowering periods support its recognition as an independent species.

We have not examined the holotype of Lepidium moabense and follow N. H. Holmgren (2005b) in reducing it to synonymy of L. eastwoodiae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (2 in the flora).

R. C. Rollins (1993) and C. L. Hitchcock (1945) divided Lepidium lasiocarpum into four and seven varieties, respectively. Hitchcock admitted that the species is highly variable in every aspect and that his varieties cover some of the “more conspicuous” variations. In our opinion, perhaps two or three groups are somewhat sufficiently defined to be recognized formally, though the lines separating them blur in some parts of the species range. We are reluctantly treating them as subspecies; subsp. palmeri (S. Watson) Thellung is restricted to Mexico (Baja California). The other infraspecific taxa recognized by those authors are based on trivial differences in leaf and indumentum, and we do not believe that they merit recognition.

The reports by R. C. Rollins (1993) and N. H. Holmgren (2005b) of four and six stamens in Lepidium lasiocarpum were most likely repeated from C. L. Hitchcock (1936, 1945b) without further verification. We have examined hundreds of specimens that definitely belong to this species, and in not a single case did we find a flower with more than two stamens. It is quite likely that the reports of more than two stamens were based on misidentified plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves hirsute (or fringed on margin), trichomes not pustular-based; fruiting pedicels 0.2-0.4(-0.6) mm wide, usually less than 3 times as wide as thick; nectar glands toothlike, to 0.2 mm.
subsp. lasiocarpum
1. Fruit valves hispid, trichomes pustular-based; fruiting pedicels (0.4-)0.5-0.7 mm wide, often more than 3 times as wide as thick; nectar glands not toothlike (subulate), 0.3-0.5 mm.
subsp. wrightii
Source FNA vol. 7, p. 581. FNA vol. 7, p. 584.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Subordinate taxa
L. lasiocarpum subsp. lasiocarpum, L. lasiocarpum subsp. wrightii
Synonyms L. alyssoides var. eastwoodiae, L. moabense, L. montanum var. eastwoodiae L. ruderale var. lasiocarpum, Nasturtium lasiocarpum
Name authority Wooton: Bull. Torrey Bot. Club 25: 258. (1898) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 115. (1838)
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