Lepidium eastwoodiae |
Lepidium integrifolium |
|
---|---|---|
mesa pepperwort |
thickleaf pepperweed |
|
Habit | Annuals, biennials, or perennials; (base woody); glabrous or pubescent. | Perennials; (caudex often thick, not aboveground, covered with persistent petiolar remains); puberulent. |
Stems | simple from base, erect, branched distally, (3.5–)4.5–16(–18) dm. |
several from base (caudex), ascending, branched distally, (1–)1.5–3.5(–4) dm. |
Basal leaves | (soon deciduous), not rosulate; petiole (1–)2–5.5(–7.5) cm; blade pinnatifid, (2–)3–6.8(–9) cm × 10–30 mm, margins (of lobes) dentate to serrate. |
rosulate; petiole (0.5–)1.5–6(–7.5) cm; blade oblanceolate to obovate, (1.5–)2.5–7(–9) cm × (10–)15–25 mm, margins usually entire, rarely denticulate subapically. |
Cauline leaves | shortly petiolate or sessile; blade narrowly lanceolate or oblanceolate to linear, 3–7 cm × (2.5–)4–10mm (smaller distally), base attenuate to cuneate, not auriculate, margins usually entire, rarely dentate. |
shortly petiolate or sessile; blade narrowly lanceolate to broadly oblanceolate, 1–5 cm × 2–9(–12) mm, base cuneate, not auriculate, margins usually entire, rarely denticulate subapically. |
Racemes | elongated in fruit; rachis puberulent, trichomes straight or curved. |
elongated in fruit; rachis puberulent, trichomes straight, sometimes clavate. |
Flowers | sepals suborbicular to oblong, 0.8–1.5 × 0.7–1.2 mm; petals white, suborbicular, 2.2–3.5(4) × 1.5–2.5 mm, claw 0.7–1.5 mm; stamens 6; filaments 1.5–2.5 mm, (glabrous); anthers 0.3–0.4 mm. |
sepals oblong-obovate, (1.5–)1.8–2.5 × 0.8–1.3 mm; petals white, obovate, (2.3)2.5–3.6(–4) × 1.5–2.2 mm, claw 0.5–1 mm; stamens (2 or) 4 (or 6), median and lateral when 4, (erect); filaments 1.7–2.5 mm, (glabrous); anthers 0.5–0.8 mm. |
Fruiting pedicels | divaricate-ascending to horizontal, usually slightly recurved or somewhat sigmoid, rarely straight, (not winged), (3–)3.5–7.5(–8) × 0.2–0.3 mm, puberulent adaxially. |
divaricate-ascending to horizontal, straight, (not winged), (4–)5–10 × 0.3–0.5 mm, puberulent adaxially. |
Fruits | broadly ovate, 2–3.5(–4) × 1.8–2.6(–3) mm, apically winged, apical notch 0.1–0.2mm deep; valves thin, smooth, not veined, glabrous; style (0.2–)0.3–0.6(–0.7) mm, exserted beyond apical notch. |
ovate, (3–)3.2–4(–4.4) × 2–3.5 mm, apically winged, apical notch 0.1–0.3 mm deep; valves thin, smooth, not veined, glabrous; style 0.5–0.8(–1) mm, exserted beyond apical notch. |
Seeds | (dark brown), ovate, 1.4–1.8 × 0.8–1.1 mm. |
ovate, 1.8–2 × 0.9–1.1 mm. |
Lepidium eastwoodiae |
Lepidium integrifolium |
|
Phenology | Flowering Jul–Sep. | Flowering Jun–Jul. |
Habitat | Pinyon-juniper, sagebrush, or mixed desert shrub communities | Alkaline and saline meadows |
Elevation | 900-2200 m (3000-7200 ft) | 1300-2000 m (4300-6600 ft) |
Distribution |
AZ; CO; NM; UT
|
AZ; NV; UT; WY |
Discussion | C. L. Hitchcock (1936) and R. C. Rollins (1993) reduced Lepidium eastwoodiae to a variety of L. montanum and L. alyssoides, respectively. However, the differences in morphology and flowering periods support its recognition as an independent species. We have not examined the holotype of Lepidium moabense and follow N. H. Holmgren (2005b) in reducing it to synonymy of L. eastwoodiae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. The circumscription of Lepidium integrifolium is somewhat controversial. C. L. Hitchcock (1936) treated it as two varieties or (Hitchcock 1950) two subspecies of L. montanum, whereas R. C. Rollins (1993) treated it as a distinct species with two varieties. Rollins indicated that the species has two stamens, but such occurrence is rather rare. Most commonly, it has four stamens and is readily distinguished from related species by having four nectar glands and sepals sparsely pubescent subapically with crisped trichomes. Nothing is known about the populational variation of stamen number in the species and whether one or more taxa are involved. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 581. | FNA vol. 7, p. 583. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | L. alyssoides var. eastwoodiae, L. moabense, L. montanum var. eastwoodiae | L. montanum subsp. integrifolium, L. montanum var. integrifolium, L. utahense, L. zionis, Nasturtium integrifolium |
Name authority | Wooton: Bull. Torrey Bot. Club 25: 258. (1898) | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 116. (1838) |
Web links |