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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop

shining pepper grass, shining pepper-weed

Habit Perennials; (rhizomatous); hirsute or glabrate. Annuals; puberulent.
Stems

often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm.

simple to numerous from base, erect to ascending or decumbent, sometimes branched distally, (0.4–)1–3.5(–4.2) dm.

Basal leaves

(early withered);

not rosulate;

petiole 1–4 cm;

blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire.

(soon withered);

not rosulate;

petiole (0.4–)1–4.2(–5) cm;

blade pinnatisect (lobes usually linear to filiform, rarely oblong to lanceolate), (0.8–)1.5–7.3(–8) cm, margins (of lobes) usually entire, rarely dentate.

Cauline leaves

sessile;

blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous).

petiolate or subsessile;

similar to basal, blade smaller, sometimes undivided and linear, base attenuate, not auriculate.

Racemes

(corymbose panicles), slightly or considerably elongated in fruit;

rachis glabrous or puberulent, trichomes straight or curved, cylindrical.

much-elongated, (lax) in fruit;

rachis puberulent to hirsutulous, trichomes straight, cylindrical.

Flowers

sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm;

petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm;

stamens 6;

filaments 2–3 mm, (glabrous);

anthers 0.4–0.5 mm.

sepals oblong-ovate, (0.7–)0.9–1.3(–1.5) × 0.5–0.8 mm;

petals (rarely rudimentary or absent), white, oblanceolate, (0.8–)1.2–2(–2.8) × 0.2–1(–1.6) mm, claw absent, (glabrous);

stamens usually 4, median, rarely 6;

filaments (0.5–)0.7–1(–1.3);

anthers 0.1–0.2 mm.

Fruiting pedicels

ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially.

usually divaricate, rarely ascending or suberect, usually slightly or weakly recurved, rarely straight, (strongly flattened), 2.5–5(–6.5) × (0.4–)0.6–0.9(–1.1) mm, puberulent adaxially and sometimes on proximal 1/3 abaxially.

Fruits

(indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent;

valves thin, smooth, reticulate-veined, glabrous;

style (0.6–)1–1.8(–2) mm.

orbicular to broadly ovate, (2.5–)3–5.5(–6.5) × (2–)2.6–5(–5.4) mm, apically winged, apical notch 0.3–0.7(–1) mm deep;

valves thin, smooth, slightly veined or not, usually glabrous, rarely minutely and sparsely puberulent (along margin);

style usually obsolete, rarely to 0.1 mm, included in apical notch.

Seeds

ovate, 1.5–2.3 × 1–1.3 mm.

ovate-oblong, 1.6–2.6 × 0.9–1.2 mm.

2n

= 32, 64.

Lepidium draba

Lepidium nitidum

Phenology Flowering Apr–Aug. Flowering Feb–Mar.
Habitat Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas Alkaline flats and sinks, meadows, pastures, dry vernal pools, fields, sandy beaches, grassy area, gravelly slopes, creosote bush desert
Elevation 0-3300 m (0-10800 ft) 0-1000 m (0-3300 ft)
Distribution
from FNA
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states.

Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

C. L. Hitchcock (1936) and R. C. Rollins (1993) recognized three varieties within Lepidium nitidum, of which var. howellii was said to differ from var. nitidum by having puberulent (versus glabrous) fruit margins and stems densely (versus glabrous or sparsely to densely) pubescent. These characters do not covary, and some glabrate plants have puberulent fruit margins, whereas some very densely pubescent plants have glabrous fruits. As for var. oreganum, it is based on plants with divergent fruit wings, but this feature shows every degree of transition from divergent to non-divergent, and both those authors accepted in var. nitidum forms with slightly divergent fruit wings. The species is highly variable in leaf division, fruit size and shape, indumentum density, and flower morphology, especially in the number of stamens and presence or absence of petals. We prefer to maintain it without any further splitting.

C. L. Hitchcock (1945b) indicated that Lepidium nitidum occurs also in Chile but did not indicate whether the plant is introduced or native there. The material that he annotated and cited as L. nitidum from Chile clearly represents misidentified L. chilense Kunze ex Walpers. The latter always has two stamens, whereas L. nitidum usually has four, rarely six, and Hitchcock’s report of two stamens in the latter species was likely based on misidentified plants of another species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 581. FNA vol. 7, p. 588.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Synonyms Cardaria draba, Cochlearia draba, Nasturtium draba L. leiocarpum, L. nitidum var. howellii, L. nitidum var. insigne, L. nitidum var. oreganum, L. oreganum, L. strictum var. oreganum, Nasturtium nitidum
Name authority Linnaeus: Sp. Pl. 2: 645. (1753) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 116. (1838)
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