Lepidium draba |
Lepidium nanum |
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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop |
dwarf pepperweed |
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Habit | Perennials; (rhizomatous); hirsute or glabrate. | Perennials; (forming pincushion-like, pulvinate mounds, caudex woody, to 1.5 cm diam., buried, much-branched, covered with persistent leaves); puberulent. |
Stems | often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm. |
simple from base (caudex branches), erect to ascending, unbranched distally, 0.05–0.2 dm. |
Basal leaves | (early withered); not rosulate; petiole 1–4 cm; blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire. |
rosulate; petiole undifferentiated; blade obovate, 2.5–5 cm × 15–25(–35) mm, margins entire, (ciliolate), apex deeply 3-lobed (lobes ovate to suborbicular, margins entire). |
Cauline leaves | sessile; blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous). |
absent. |
Racemes | (corymbose panicles), slightly or considerably elongated in fruit; rachis glabrous or puberulent, trichomes straight or curved, cylindrical. |
slightly elongated in fruit, (2–7-fruited); rachis puberulent, trichomes straight, cylindrical. |
Flowers | sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm; petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm; stamens 6; filaments 2–3 mm, (glabrous); anthers 0.4–0.5 mm. |
sepals (tardily deciduous), obovate, 1.3–4 × 0.8–1.1 mm; petals pale yellow or creamy white, spatulate, 1.8–2.9 × 0.8–1.2 mm, claw 0.8–1.1 mm; stamens 6; filaments 1.4–2 mm, (glabrous); anthers 1.4–2 mm. |
Fruiting pedicels | ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially. |
suberect to ascending, often straight, (terete), 2–4.5 × 0.2–0.3 mm, puberulent throughout. |
Fruits | (indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent; valves thin, smooth, reticulate-veined, glabrous; style (0.6–)1–1.8(–2) mm. |
ovate, 2–4.2 × 1.5–3 mm, often apically winged, apical notch 0.1–0.2 mm deep; valves thin, smooth, not veined, glabrous; style (0.4–)0.6–1(–1.2) mm, exserted beyond apical notch. |
Seeds | ovate, 1.5–2.3 × 1–1.3 mm. |
oblong, 1–2 × 0.8–1 mm. |
2n | = 32, 64. |
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Lepidium draba |
Lepidium nanum |
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Phenology | Flowering Apr–Aug. | Flowering May–Jun. |
Habitat | Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas | Gypsum knolls, tufa mounds around hotsprings, quartzite gravel, barren areas with shale and chalky soil, gravelly hillsides, white calcareous soils |
Elevation | 0-3300 m (0-10800 ft) | 1500-2200 m (4900-7200 ft) |
Distribution |
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
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NV; UT |
Discussion | Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states. Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Lepidium nanum is most common in Nevada and is known in Utah from collections in Tooele County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 581. | FNA vol. 7, p. 587. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | Cardaria draba, Cochlearia draba, Nasturtium draba | |
Name authority | Linnaeus: Sp. Pl. 2: 645. (1753) | S. Watson: Botany (Fortieth Parallel), 30, plate 4, figs. 5–7. (1871) |
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