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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop

dwarf pepper grass, San Diego pepperweed

Habit Perennials; (rhizomatous); hirsute or glabrate. Annuals; puberulent or hirsute.
Stems

often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm.

simple or several from base, erect to ascending or (outer ones) decumbent, unbranched or branched, 0.2–1.5(–3.8) dm.

Basal leaves

(early withered);

not rosulate;

petiole 1–4 cm;

blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire.

(soon withered);

not rosulate;

petiole often undifferentiated (to 3 cm);

blade linear, 2–10 cm × 1–4 mm, margins entire, dentate, or pinnatisect (lobes 2–10 pairs, margins entire or dentate).

Cauline leaves

sessile;

blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous).

similar to basal, smaller, blade base attenuate, not auriculate, margins entire.

Racemes

(corymbose panicles), slightly or considerably elongated in fruit;

rachis glabrous or puberulent, trichomes straight or curved, cylindrical.

(subcapitate to cylindrical), elongated or not in fruit, (compact);

rachis puberulent, trichomes straight, cylindrical.

Flowers

sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm;

petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm;

stamens 6;

filaments 2–3 mm, (glabrous);

anthers 0.4–0.5 mm.

sepals (somewhat persistent), ovate, 1.1–1.4 × 0.6–0.8 mm;

petals greenish, obovate-oblong, 1.9–3 × 0.8–1.3 mm, claw absent, (usually pubescent outside, with fringed margin, rarely glabrescent);

stamens 4, median;

filaments 0.8–1.1 mm;

anthers 0.15–0.2 mm.

Fruiting pedicels

ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially.

erect to slightly ascending, straight and appressed to rachis or distally slightly recurved, (strongly flattened), 2.5–5 × 0.9–1.4 mm, usually puberulent throughout, rarely only adaxially.

Fruits

(indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent;

valves thin, smooth, reticulate-veined, glabrous;

style (0.6–)1–1.8(–2) mm.

oblong-ovate, 5–7 × 2.8–4 mm, apically winged, apical notch 1.4–2.8 mm deep;

valves thick, smooth, strongly reticulate-veined, hirsute and puberulent, (trichomes spreading, mixed with smaller ones);

style obsolete, included in apical notch.

Seeds

ovate, 1.5–2.3 × 1–1.3 mm.

oblong, 2–2.4 × 1.1–1.3 mm.

2n

= 32, 64.

Lepidium draba

Lepidium latipes

Phenology Flowering Apr–Aug. Flowering Mar–Jun.
Habitat Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas Margins of vernal pools, edges of salt marshes, alkaline flats and adobe, pastures, mud-wet fields
Elevation 0-3300 m (0-10800 ft) 0-700 m (0-2300 ft)
Distribution
from FNA
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states.

Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variety heckardii, which is said to differ from var. latipes mainly by having elongated stems simple at base (instead of short and branched basally), grows mixed with var. latipes in single populations. It appears that the difference is trivial and may well be controlled by a few-gene difference. In our opinion, formal distinction is unwarranted; similar conditions exist in other species (e.g., 27. Lepidium nitidum).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 581. FNA vol. 7, p. 585.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Synonyms Cardaria draba, Cochlearia draba, Nasturtium draba L. latipes var. heckardii, Nasturtium latipes
Name authority Linnaeus: Sp. Pl. 2: 645. (1753) Hooker: Icon. Pl. 1: plate 41. (1836)
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