Lepidium draba |
Lepidium integrifolium |
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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop |
thickleaf pepperweed |
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Habit | Perennials; (rhizomatous); hirsute or glabrate. | Perennials; (caudex often thick, not aboveground, covered with persistent petiolar remains); puberulent. |
Stems | often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm. |
several from base (caudex), ascending, branched distally, (1–)1.5–3.5(–4) dm. |
Basal leaves | (early withered); not rosulate; petiole 1–4 cm; blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire. |
rosulate; petiole (0.5–)1.5–6(–7.5) cm; blade oblanceolate to obovate, (1.5–)2.5–7(–9) cm × (10–)15–25 mm, margins usually entire, rarely denticulate subapically. |
Cauline leaves | sessile; blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous). |
shortly petiolate or sessile; blade narrowly lanceolate to broadly oblanceolate, 1–5 cm × 2–9(–12) mm, base cuneate, not auriculate, margins usually entire, rarely denticulate subapically. |
Racemes | (corymbose panicles), slightly or considerably elongated in fruit; rachis glabrous or puberulent, trichomes straight or curved, cylindrical. |
elongated in fruit; rachis puberulent, trichomes straight, sometimes clavate. |
Flowers | sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm; petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm; stamens 6; filaments 2–3 mm, (glabrous); anthers 0.4–0.5 mm. |
sepals oblong-obovate, (1.5–)1.8–2.5 × 0.8–1.3 mm; petals white, obovate, (2.3)2.5–3.6(–4) × 1.5–2.2 mm, claw 0.5–1 mm; stamens (2 or) 4 (or 6), median and lateral when 4, (erect); filaments 1.7–2.5 mm, (glabrous); anthers 0.5–0.8 mm. |
Fruiting pedicels | ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially. |
divaricate-ascending to horizontal, straight, (not winged), (4–)5–10 × 0.3–0.5 mm, puberulent adaxially. |
Fruits | (indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent; valves thin, smooth, reticulate-veined, glabrous; style (0.6–)1–1.8(–2) mm. |
ovate, (3–)3.2–4(–4.4) × 2–3.5 mm, apically winged, apical notch 0.1–0.3 mm deep; valves thin, smooth, not veined, glabrous; style 0.5–0.8(–1) mm, exserted beyond apical notch. |
Seeds | ovate, 1.5–2.3 × 1–1.3 mm. |
ovate, 1.8–2 × 0.9–1.1 mm. |
2n | = 32, 64. |
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Lepidium draba |
Lepidium integrifolium |
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Phenology | Flowering Apr–Aug. | Flowering Jun–Jul. |
Habitat | Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas | Alkaline and saline meadows |
Elevation | 0-3300 m (0-10800 ft) | 1300-2000 m (4300-6600 ft) |
Distribution |
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
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AZ; NV; UT; WY |
Discussion | Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states. Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. The circumscription of Lepidium integrifolium is somewhat controversial. C. L. Hitchcock (1936) treated it as two varieties or (Hitchcock 1950) two subspecies of L. montanum, whereas R. C. Rollins (1993) treated it as a distinct species with two varieties. Rollins indicated that the species has two stamens, but such occurrence is rather rare. Most commonly, it has four stamens and is readily distinguished from related species by having four nectar glands and sepals sparsely pubescent subapically with crisped trichomes. Nothing is known about the populational variation of stamen number in the species and whether one or more taxa are involved. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 581. | FNA vol. 7, p. 583. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | Cardaria draba, Cochlearia draba, Nasturtium draba | L. montanum subsp. integrifolium, L. montanum var. integrifolium, L. utahense, L. zionis, Nasturtium integrifolium |
Name authority | Linnaeus: Sp. Pl. 2: 645. (1753) | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 116. (1838) |
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