Lepidium draba |
Lepidium alyssoides |
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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop |
mesa pepperwort |
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Habit | Perennials; (rhizomatous); hirsute or glabrate. | Perennials or subshrubs; (woody base often aboveground); glabrous or minutely puberulent. |
Stems | often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm. |
few to several from base, erect to ascending, branched throughout, (0.7–)1–4.8(–6.1) dm. |
Basal leaves | (early withered); not rosulate; petiole 1–4 cm; blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire. |
often not rosulate; petiole 1–6 cm; blade pinnately lobed, (1–)1.5–8(–11) cm × (5–)10–35 mm, margins (of lobes) entire or denticulate. |
Cauline leaves | sessile; blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous). |
sessile; blade linear, (0.8–)1.3–7(–9.5) cm × (0.7–)1–2(–3) mm, base attenuate, not auriculate, margins entire. |
Racemes | (corymbose panicles), slightly or considerably elongated in fruit; rachis glabrous or puberulent, trichomes straight or curved, cylindrical. |
elongated in fruit; rachis puberulent or glabrous. |
Flowers | sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm; petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm; stamens 6; filaments 2–3 mm, (glabrous); anthers 0.4–0.5 mm. |
sepals ovate to oblong, 1–2 × 0.8–1 mm; petals white, suborbicular, 2–3 × 1–2 mm, claw 0.5–1.5 mm; stamens 6; filaments 1.5–2 mm, (glabrous); anthers 0.2–0.4 mm. |
Fruiting pedicels | ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially. |
divaricate to horizontal, straight or recurved to somewhat sigmoid, (terete), 3.5–8(–11) × 0.2 mm, glabrous or puberulent adaxially. |
Fruits | (indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent; valves thin, smooth, reticulate-veined, glabrous; style (0.6–)1–1.8(–2) mm. |
broadly ovate, 2–3.7(–4.3) × (1.5–)1.8–2.9(–3.4) mm, apically winged, apical notch 0.1–0.3(–0.4) mm deep; valves thin, smooth, not veined, glabrous; style 0.2–0.6 mm, exserted beyond apical notch. |
Seeds | ovate, 1.5–2.3 × 1–1.3 mm. |
ovate, 1.5–1.8(–2) × 0.9–1.2(–1.5) mm. |
2n | = 32, 64. |
= 32. |
Lepidium draba |
Lepidium alyssoides |
|
Phenology | Flowering Apr–Aug. | Flowering May–Jul. |
Habitat | Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas | Pinyon-juniper or sagebrush communities, prairies, grasslands, sandstone outcrops, gypsum flats, sand dunes, dry flats and river bottoms, gravelly roadsides |
Elevation | 0-3300 m (0-10800 ft) | 1200-2800 m (3900-9200 ft) |
Distribution |
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
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AZ; CO; NM; NV; TX; UT; WY; Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí)
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Discussion | Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states. Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of the five varieties of Lepidium alyssoides recognized by R. C. Rollins (1993), one (var. mexicanum Rollins) is a short-tufted form of the species restricted to Mexico that does not seem to merit recognition, another (var. junceum) is a glabrescent form of the type variety, a third (var. eastwoodiae) is treated below as a distinct species, and the fourth (var. angustifolium) is included here within L. alyssoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 581. | FNA vol. 7, p. 575. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | Cardaria draba, Cochlearia draba, Nasturtium draba | L. alyssoides var. angustifolium, L. alyssoides var. junceum, L. alyssoides var. minus, L. alyssoides var. polycarpum, L. alyssoides var. streptocarpum, L. montanum subsp. alyssoides, L. montanum var. alyssoides, L. montanum subsp. angustifolium, L. montanum var. angustifolium, L. tortum |
Name authority | Linnaeus: Sp. Pl. 2: 645. (1753) | A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 10. (1849) |
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