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heart-pod hoary-cress, heart-pod pepperweed, hoary cress, hoary pepperwort, whitetop

alkali pepperwort, alkali veiny pepper-grass, net pepper grass

Habit Perennials; (rhizomatous); hirsute or glabrate. Annuals; hirsute or puberulent.
Stems

often simple from base, erect or decumbent basally, branched (several) distally, (0.8–)2–6.5(–9) dm.

few to several from base, erect to ascending, unbranched, (0.5–)0.8–3 dm.

Basal leaves

(early withered);

not rosulate;

petiole 1–4 cm;

blade obovate, spatulate, or ovate, (1.5–)3–10(–15) cm × 10–40 mm, margins sinuate to dentate or entire.

(soon withered);

not rosulate;

petiole 1–4 cm;

blade linear or pinnatisect, (2–)2.5–6.1(–7.2) cm × (0.5–)1–2(–3) mm, (lobes linear to narrowly oblong), margins entire.

Cauline leaves

sessile;

blade ovate, elliptic, oblong, or lanceolate, oblanceolate, or obovate, (1–)3–9(–15) cm × (5–)10–20(–50) mm, base sagittate-amplexicaul or auriculate, margins dentate or entire, (surfaces pubescent or glabrous).

sessile;

blade linear, 1.2–5.8 cm × 0.5–2(–3) mm, base attenuate, not auriculate, margins entire.

Racemes

(corymbose panicles), slightly or considerably elongated in fruit;

rachis glabrous or puberulent, trichomes straight or curved, cylindrical.

elongated, (dense or lax) in fruit;

rachis puberulent or hirsute, trichomes straight, cylindrical.

Flowers

sepals oblong to ovate, 1.5–2.5 × 0.7–1.2 mm;

petals white, obovate, (2.5–)3–4(–4.5) × (1–)1.3–2(–2.2) mm, claw 1–1.7 mm;

stamens 6;

filaments 2–3 mm, (glabrous);

anthers 0.4–0.5 mm.

sepals oblong to ovate, 0.7–1.1 × 0.2–0.6 mm;

petals absent;

stamens 4, median;

filaments 0.5–1 mm;

anthers ca. 0.1 mm.

Fruiting pedicels

ascending to horizontal, straight, (terete), 5–10(–15) × 0.2–0.3 mm, glabrous or sparsely puberulent adaxially.

erect to slightly ascending, straight and appressed to rachis or distally slightly recurved, (strongly flattened), (2–)3–4.2 × 0.4–0.6 mm (width proximal to apex), puberulent throughout.

Fruits

(indehiscent), cordate to subreniform, (2–)2.5–3.7(–4.3) × (3.2–)3.7–5(–5.6) mm, apically (obtuse to subacute), not winged, apical notch absent;

valves thin, smooth, reticulate-veined, glabrous;

style (0.6–)1–1.8(–2) mm.

ovate to ovate-oblong, (3–)4–6 × 2.5–3.5 mm, apically winged, apical notch (V-shaped), (0.8–)1–2 mm deep;

valves thin, smooth, strongly reticulate-veined, usually glabrous, rarely pubescent;

style absent or obsolete, included in apical notch.

Seeds

ovate, 1.5–2.3 × 1–1.3 mm.

ovate, 1.3–2 × 0.8–1.2 mm.

2n

= 32, 64.

Lepidium draba

Lepidium acutidens

Phenology Flowering Apr–Aug. Flowering Feb–Apr.
Habitat Mountain slopes, roadsides, fields, agricultural lands, stream sides, disturbed grounds, pastures, waste areas Alkaline flats, gullies, or fields, saline vernal flats, grassy fields
Elevation 0-3300 m (0-10800 ft) 0-400 m (0-1300 ft)
Distribution
from FNA
AZ; CA; CO; CT; DC; IA; ID; IL; IN; KS; MA; MD; MI; MN; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SD; UT; WA; WY; AB; BC; MB; NS; ON; SK; s Europe; sw Asia [Introduced in North America; introduced also in Mexico (Distrito Federal), South America, s Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Lepidium draba is poorly established and known from old collections in the eastern part of the United States, it has become a noxious weed in several western states.

Lepidium draba and its nearest relatives, L. appelianum and L. chalepense, form a monophyletic clade most closely related to L. campestre (K. Mummenhoff et al. 2001). A. Thellung (1906) and C. L. Hitchcock (1936) correctly placed L. draba in Lepidium, as did Linnaeus. The recognition of the first three species in Cardaria and the maintenance of their nearest relative, L. campestre, in Lepidium do not make any sense on both phylogenetic and taxonomic grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Lepidium acutidens was treated by C. L. Hitchcock (1936) and R. C. Rollins (1993) as a variety of L. dictyotum. The differences in the fruits as well as the absence of intermediates between them, despite the overlap of their ranges in California, justify their recognition as independent species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 581. FNA vol. 7, p. 575.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Synonyms Cardaria draba, Cochlearia draba, Nasturtium draba L. dictyotum var. acutidens, L. oxycarpum var. acutidens
Name authority Linnaeus: Sp. Pl. 2: 645. (1753) (A. Gray) Howell: Fl. N.W. Amer., 64. (1897)
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