Lepidium dictyotum |
Lepidium tiehmii |
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alkali pepper-weed, alkali peppergrass, alkali pepperwort, net pepper grass, veiny pepper-grass |
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Habit | Annuals; hirsute. | Perennials; (caudex woody, to 1 cm diam., covered with persistent petiolar remains); glabrous throughout. |
Stems | few to several from base, erect to ascending, or (outer ones) decumbent, unbranched, (0.15–)0.3–1.3(–2.1) dm. |
simple from base (caudex branch), erect, branched distally, 1–7.5 dm. |
Basal leaves | (soon withered); not rosulate; petiole 0.5–2 cm; blade pinnatifid to pinnatisect (lobes linear to narrowly oblong), (1.5–)2.2–5.7(–7) cm × (0.5–)1–2(–3) mm, margins entire. |
rosulate; petiole (1.5–)2.5–10(–13) cm; blade oblong to lanceolate, (2.5–)4.5–9(–14) cm × 15–40 mm, margins entire. |
Cauline leaves | sessile; blade usually linear, rarely with linear lobes, 1–5 cm × 0.5–2 mm, base attenuate, not auriculate, margins entire. |
shortly petiolate; blade oblanceolate, (much smaller than basal), base attenuate-cuneate, not auriculate, margins entire. |
Racemes | elongated, (dense) in fruit; rachis hirsute, trichomes straight, cylindrical. |
(paniculate), considerably elongated in fruit. |
Flowers | sepals oblong to ovate, 0.7–1.1 × 0.2–0.6 mm; petals absent; stamens 4, median; filaments 0.5–1 mm; anthers ca. 0.1 mm. |
sepals oblong, 2.5–3(–4) × 1.2–1.8 mm; petals creamy white to pale yellow, obovate to oblanceolate, 4–5.5(–6.5) × 2–3.3 mm, claw 1.5–2.2 mm; stamens 6; filaments (median pairs) 3.5–4.5 mm; anthers 1–1.2 mm. |
Fruiting pedicels | erect to slightly ascending, straight and appressed to rachis or distally slightly recurved, (strongly flattened), (1.6–)1.9–2.5(–3) × 0.4–0.8 mm (width proximal to apex), usually hirsute or, rarely, only adaxially. |
divaricate-ascending, straight, (terete), 8–15 × 0.4–0.5 mm. |
Fruits | ovate, 2.5–3.5 × 2–2.8 mm, apically winged, apical notch (closed, often U-shaped), 0.5–0.7(–0.8) mm deep; valves thin, smooth, strongly reticulate-veined, usually hirsute, rarely glabrous; style absent or obsolete, included in apical notch. |
obovate to somewhat rhomboid, 7–11 × 5–6.5 mm, apically not winged, apical notch absent; valves thin, smooth, obscurely veined; style 0.2–0.6 mm. |
Seeds | ovate, 1.2–1.8 × 0.8–1.2 mm. |
oblong, 3.7–4.5 × 1.6–2 mm. |
Lepidium dictyotum |
Lepidium tiehmii |
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Phenology | Flowering Mar–Jun. | Flowering Apr–Jun. |
Habitat | Margins of playas, saline areas, meadows, gypsum hills, dried pools, alkaline and clay flats and sinks, near hot springs, roadsides, borders of springs and ponds, sandy flats | Rocky crevices and slopes in sagebrush communities |
Elevation | 0-1600 m (0-5200 ft) | 1400-1800 m (4600-5900 ft) |
Distribution |
CA; ID; NV; OR; UT; WA; Mexico (Baja California)
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NV |
Discussion | Of conservation concern. Lepidium tiehmii is known from mountain ranges in Douglas and Lyon counties. It was described and has been maintained (N. H. Holmgren 2005b) in Stroganowia, a genus now united with Lepidium (I. A. Al-Shehbaz et al. 2002) that otherwise is disjunct and restricted to the central Asian states of the Former Soviet Union and adjacent western China. In our opinion, the similarity of this species to those Asian ones formerly placed in Stroganowia is superficial and is the result of convergence rather than descent. The cotyledonary type was erroneously reported as conduplicate (R. C. Rollins 1993; N. H. Holmgren 2005b). In the several seeds that we dissected it was always incumbent. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 580. | FNA vol. 7, p. 594. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | L. acutidens var. microcarpum, L. dictyotum var. macrocarpum, Nasturtium dictyotum | Stroganowia tiehmii |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 329. (1868) | (Rollins) Al-Shehbaz: Novon 12: 9. (2002) |
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