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alkali pepper-weed, alkali peppergrass, alkali pepperwort, net pepper grass, veiny pepper-grass

hairypod pepperweed, shaggyfruit pepperweed

Habit Annuals; hirsute. Annuals; hirsute or hispid, (trichomes cylindrical).
Stems

few to several from base, erect to ascending, or (outer ones) decumbent, unbranched, (0.15–)0.3–1.3(–2.1) dm.

usually few to several, rarely simple from base, erect to ascending or (outer ones) decumbent, branched distally, (0.15–)0.6–3(–3.8) dm.

Basal leaves

(soon withered);

not rosulate;

petiole 0.5–2 cm;

blade pinnatifid to pinnatisect (lobes linear to narrowly oblong), (1.5–)2.2–5.7(–7) cm × (0.5–)1–2(–3) mm, margins entire.

(later withered);

not rosulate;

petiole (0.4–)1–3.5(–5) cm;

blade spatulate to oblanceolate, lyrate-pinnatifid, pinnatisect, or 2-pinnatifid, (0.7–)1.5–4.5(–7.5) cm × (9–)12–20(–30) mm, margins rarely dentate, (lobes) entire or dentate.

Cauline leaves

sessile;

blade usually linear, rarely with linear lobes, 1–5 cm × 0.5–2 mm, base attenuate, not auriculate, margins entire.

subsessile or petioles 0.8–2.2 cm, blade lanceolate to oblanceolate, (0.7–)1.2–3.3(–5) cm × (2–)4–12 mm, base cuneate, not auriculate, margins subentire to dentate.

Racemes

elongated, (dense) in fruit;

rachis hirsute, trichomes straight, cylindrical.

often considerably elongated in fruit;

rachis hirsute or hispid, trichomes straight, cylindrical.

Flowers

sepals oblong to ovate, 0.7–1.1 × 0.2–0.6 mm;

petals absent;

stamens 4, median;

filaments 0.5–1 mm;

anthers ca. 0.1 mm.

sepals oblong, 1–1.3(–1.5) × 0.5–0.8mm;

petals (sometimes absent), white, oblanceolate to linear, (0.3–)0.6–1.5(–2) × (0.1–)0.2–0.5 mm, claw absent;

stamens 2, median;

filaments 1–1.4 mm;

anthers 0.2–0.3 mm.

Fruiting pedicels

erect to slightly ascending, straight and appressed to rachis or distally slightly recurved, (strongly flattened), (1.6–)1.9–2.5(–3) × 0.4–0.8 mm (width proximal to apex), usually hirsute or, rarely, only adaxially.

divaricate-ascending to horizontal, straight or slightly curved, (often strongly flattened), (1.8–)2–4(–4.6) × 0.2–0.7 mm (to 0.3 mm thick), hirsute to hispid throughout or adaxially.

Fruits

ovate, 2.5–3.5 × 2–2.8 mm, apically winged, apical notch (closed, often U-shaped), 0.5–0.7(–0.8) mm deep;

valves thin, smooth, strongly reticulate-veined, usually hirsute, rarely glabrous;

style absent or obsolete, included in apical notch.

ovate to ovate-orbicular, 2.8–4(–4.6) × 2.4–3.6(–4) mm, (base broadly cuneate to rounded), apically winged, apical notch (0.2–)0.3–0.6(–0.7) mm deep;

valves thin, smooth, not veined, hirsute to hispid (on surface or margin);

style obsolete or to 0.1 mm, included in apical notch.

Seeds

ovate, 1.2–1.8 × 0.8–1.2 mm.

ovate, 1.4–2.2 × 0.9–1.4 mm.

Lepidium dictyotum

Lepidium lasiocarpum

Phenology Flowering Mar–Jun.
Habitat Margins of playas, saline areas, meadows, gypsum hills, dried pools, alkaline and clay flats and sinks, near hot springs, roadsides, borders of springs and ponds, sandy flats
Elevation 0-1600 m (0-5200 ft)
Distribution
from FNA
CA; ID; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; NV; TX; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (2 in the flora).

R. C. Rollins (1993) and C. L. Hitchcock (1945) divided Lepidium lasiocarpum into four and seven varieties, respectively. Hitchcock admitted that the species is highly variable in every aspect and that his varieties cover some of the “more conspicuous” variations. In our opinion, perhaps two or three groups are somewhat sufficiently defined to be recognized formally, though the lines separating them blur in some parts of the species range. We are reluctantly treating them as subspecies; subsp. palmeri (S. Watson) Thellung is restricted to Mexico (Baja California). The other infraspecific taxa recognized by those authors are based on trivial differences in leaf and indumentum, and we do not believe that they merit recognition.

The reports by R. C. Rollins (1993) and N. H. Holmgren (2005b) of four and six stamens in Lepidium lasiocarpum were most likely repeated from C. L. Hitchcock (1936, 1945b) without further verification. We have examined hundreds of specimens that definitely belong to this species, and in not a single case did we find a flower with more than two stamens. It is quite likely that the reports of more than two stamens were based on misidentified plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves hirsute (or fringed on margin), trichomes not pustular-based; fruiting pedicels 0.2-0.4(-0.6) mm wide, usually less than 3 times as wide as thick; nectar glands toothlike, to 0.2 mm.
subsp. lasiocarpum
1. Fruit valves hispid, trichomes pustular-based; fruiting pedicels (0.4-)0.5-0.7 mm wide, often more than 3 times as wide as thick; nectar glands not toothlike (subulate), 0.3-0.5 mm.
subsp. wrightii
Source FNA vol. 7, p. 580. FNA vol. 7, p. 584.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Subordinate taxa
L. lasiocarpum subsp. lasiocarpum, L. lasiocarpum subsp. wrightii
Synonyms L. acutidens var. microcarpum, L. dictyotum var. macrocarpum, Nasturtium dictyotum L. ruderale var. lasiocarpum, Nasturtium lasiocarpum
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 329. (1868) Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 115. (1838)
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