Lepidium dictyotum |
Lepidium lasiocarpum |
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alkali pepper-weed, alkali peppergrass, alkali pepperwort, net pepper grass, veiny pepper-grass |
hairypod pepperweed, shaggyfruit pepperweed |
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Habit | Annuals; hirsute. | Annuals; hirsute or hispid, (trichomes cylindrical). | ||||
Stems | few to several from base, erect to ascending, or (outer ones) decumbent, unbranched, (0.15–)0.3–1.3(–2.1) dm. |
usually few to several, rarely simple from base, erect to ascending or (outer ones) decumbent, branched distally, (0.15–)0.6–3(–3.8) dm. |
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Basal leaves | (soon withered); not rosulate; petiole 0.5–2 cm; blade pinnatifid to pinnatisect (lobes linear to narrowly oblong), (1.5–)2.2–5.7(–7) cm × (0.5–)1–2(–3) mm, margins entire. |
(later withered); not rosulate; petiole (0.4–)1–3.5(–5) cm; blade spatulate to oblanceolate, lyrate-pinnatifid, pinnatisect, or 2-pinnatifid, (0.7–)1.5–4.5(–7.5) cm × (9–)12–20(–30) mm, margins rarely dentate, (lobes) entire or dentate. |
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Cauline leaves | sessile; blade usually linear, rarely with linear lobes, 1–5 cm × 0.5–2 mm, base attenuate, not auriculate, margins entire. |
subsessile or petioles 0.8–2.2 cm, blade lanceolate to oblanceolate, (0.7–)1.2–3.3(–5) cm × (2–)4–12 mm, base cuneate, not auriculate, margins subentire to dentate. |
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Racemes | elongated, (dense) in fruit; rachis hirsute, trichomes straight, cylindrical. |
often considerably elongated in fruit; rachis hirsute or hispid, trichomes straight, cylindrical. |
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Flowers | sepals oblong to ovate, 0.7–1.1 × 0.2–0.6 mm; petals absent; stamens 4, median; filaments 0.5–1 mm; anthers ca. 0.1 mm. |
sepals oblong, 1–1.3(–1.5) × 0.5–0.8mm; petals (sometimes absent), white, oblanceolate to linear, (0.3–)0.6–1.5(–2) × (0.1–)0.2–0.5 mm, claw absent; stamens 2, median; filaments 1–1.4 mm; anthers 0.2–0.3 mm. |
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Fruiting pedicels | erect to slightly ascending, straight and appressed to rachis or distally slightly recurved, (strongly flattened), (1.6–)1.9–2.5(–3) × 0.4–0.8 mm (width proximal to apex), usually hirsute or, rarely, only adaxially. |
divaricate-ascending to horizontal, straight or slightly curved, (often strongly flattened), (1.8–)2–4(–4.6) × 0.2–0.7 mm (to 0.3 mm thick), hirsute to hispid throughout or adaxially. |
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Fruits | ovate, 2.5–3.5 × 2–2.8 mm, apically winged, apical notch (closed, often U-shaped), 0.5–0.7(–0.8) mm deep; valves thin, smooth, strongly reticulate-veined, usually hirsute, rarely glabrous; style absent or obsolete, included in apical notch. |
ovate to ovate-orbicular, 2.8–4(–4.6) × 2.4–3.6(–4) mm, (base broadly cuneate to rounded), apically winged, apical notch (0.2–)0.3–0.6(–0.7) mm deep; valves thin, smooth, not veined, hirsute to hispid (on surface or margin); style obsolete or to 0.1 mm, included in apical notch. |
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Seeds | ovate, 1.2–1.8 × 0.8–1.2 mm. |
ovate, 1.4–2.2 × 0.9–1.4 mm. |
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Lepidium dictyotum |
Lepidium lasiocarpum |
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Phenology | Flowering Mar–Jun. | |||||
Habitat | Margins of playas, saline areas, meadows, gypsum hills, dried pools, alkaline and clay flats and sinks, near hot springs, roadsides, borders of springs and ponds, sandy flats | |||||
Elevation | 0-1600 m (0-5200 ft) | |||||
Distribution |
CA; ID; NV; OR; UT; WA; Mexico (Baja California)
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AZ; CA; CO; NM; NV; TX; UT; n Mexico
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Discussion | Subspecies 3 (2 in the flora). R. C. Rollins (1993) and C. L. Hitchcock (1945) divided Lepidium lasiocarpum into four and seven varieties, respectively. Hitchcock admitted that the species is highly variable in every aspect and that his varieties cover some of the “more conspicuous” variations. In our opinion, perhaps two or three groups are somewhat sufficiently defined to be recognized formally, though the lines separating them blur in some parts of the species range. We are reluctantly treating them as subspecies; subsp. palmeri (S. Watson) Thellung is restricted to Mexico (Baja California). The other infraspecific taxa recognized by those authors are based on trivial differences in leaf and indumentum, and we do not believe that they merit recognition. The reports by R. C. Rollins (1993) and N. H. Holmgren (2005b) of four and six stamens in Lepidium lasiocarpum were most likely repeated from C. L. Hitchcock (1936, 1945b) without further verification. We have examined hundreds of specimens that definitely belong to this species, and in not a single case did we find a flower with more than two stamens. It is quite likely that the reports of more than two stamens were based on misidentified plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 580. | FNA vol. 7, p. 584. | ||||
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | L. acutidens var. microcarpum, L. dictyotum var. macrocarpum, Nasturtium dictyotum | L. ruderale var. lasiocarpum, Nasturtium lasiocarpum | ||||
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 329. (1868) | Nuttall: in J. Torrey and A. Gray, Fl. N. Amer. 1: 115. (1838) | ||||
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