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baby-seed pepper-weed, common pepper-grass, common pepper-weed, elongate peppergrass, hairy-fruit peppergrass, large-fruit peppergrass, miner's pepper-weed, prairie pepper-grass, prairie pepperweed

field cress, field pepper-grass, field pepper-weed, field peppergrass pepperwort

Habit Annuals or biennials; puberulent or glabrous, (trichomes cylindrical). Annuals; densely hirsute.
Stems

simple from base, erect, branched distally, (1–)2.5–5(–6.5) dm.

simple from base, erect, unbranched or branched distally, (0.8–)1.2–5(–6.3) dm.

Basal leaves

(early withered); rosulate;

petiole 0.5–1.5(–2) cm;

blade oblanceolate, spatulate, or oblong, (1.5–)2.5–8(–11) cm × 5–10(–20) mm, margins coarsely serrate or pinnatifid.

rosulate;

petiole (0.5–)1.5–6 cm;

blade oblanceolate or oblong, (1–)2–6(–8) cm × 5–15 mm, margins entire, lyrate, or pinnatifid.

Cauline leaves

shortly petiolate;

blade narrowly oblanceolate or linear, (0.7–)1.3–6.2(–8) cm × (0.5–)1.5–10(–18) mm, base attenuate to cuneate, not auriculate, margins usually entire or irregularly serrate to dentate, rarely pinnatifid.

sessile; oblong, lanceolate, or narrowly deltate-lanceolate, (0.7–)1–4(–6.5) cm × (2–)5–10(–15) mm, base sagittate or auriculate, margins dentate or subentire.

Racemes

considerably elongated in fruit;

rachis puberulent, trichomes straight, slender to subclavate.

much-elongated in fruit;

rachis hirsute, trichomes spreading, straight.

Flowers

sepals oblong, 0.5–0.8(–1) × 0.3–0.5 mm;

petals (absent or rudimentary) white, filiform, 0.3–0.9 mm, claw absent;

stamens 2, median;

filaments 0.6–1(–1.8) mm;

anthers 0.1–0.2 mm.

sepals oblong, (1–)1.3–1.8 × 0.6–0.8 mm;

petals white, spatulate, (1.5–)1.8–2.5(–3) × (0.2–)0.5–0.7 mm, claw 0.6–1 mm;

stamens 6;

filaments (1.2–)1.5–1.8(–2) mm, (glabrous);

anthers 0.3–0.5 mm.

Fruiting pedicels

divaricate-ascending to horizontal, straight or slightly recurved, (terete), (1.5–)2–3.5(–4) × 0.15–0.25 mm, puberulent adaxially.

horizontal, straight or slightly recurved, (terete), (3–)4–8(–10) × 0.3–0.4 mm, hirsute.

Fruits

obovate to obovate-suborbicular, (2–)2.5–3(–3.5) × 1.5–2.5(–3) mm (widest beyond middle), apically winged, apical notch 0.2–0.4 mm deep;

valves thin, smooth, not veined, glabrous or sparsely puberulent (at least on margin);

style 0.1–0.2 mm, included in apical notch.

broadly oblong to ovate, (4–)5–6(–6.5) × (3–)4–5 mm, (curved adaxially), apically broadly winged, apical notch (0.2–)0.4–0.6 mm deep;

valves thin, papillate except for wing, not veined;

style 0.2–0.5(–0.7) mm, slightly exserted beyond, or included in, apical notch.

Seeds

ovate, 1.1–1.3(–1.5) × 0.8–0.9 mm.

(dark brown), ovoid, 2–2.3(–2.8) × 1–1.4 mm.

2n

= 32.

= 16.

Lepidium densiflorum

Lepidium campestre

Phenology Flowering May–Jul. Flowering May–Jun.
Habitat Waste places, disturbed sites, prairies, fields, pastures, grasslands, chaparral, meadows, sagebrush flats, flood plains, gravelly hillsides, rock crevices, seashores, sandy places, shaley barrens, open mesa, roadsides Roadside, pastures, gardens, open flats, pine woodlands, rocky slopes, forests, waste grounds, disturbed areas, meadows, fields
Elevation 0-3500 m (0-11500 ft) 0-2600 m (0-8500 ft)
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Mexico (Chihuahua, Nuevo León) [Introduced in Europe, Asia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; WY; BC; NF; NS; ON; PE; QC; SK; Europe; Asia [Introduced in North America; introduced also in South America, South Africa]
[WildflowerSearch map]
[BONAP county map]
Discussion

North American records of Lepidium apetalum Willdenow mostly represent misidentifications of L. densiflorum. The latter has obovate fruits widest beyond the middle, whereas L. apetalum has elliptic fruits widest at the middle.

The number and limits of the varieties recognized in Lepidium densiflorum, as well as the characters used to delimit them, vary among authors (A. Thellung 1906; C. L. Hitchcock 1936; G. A. Mulligan 1961; R. C. Rollins 1993; N. H. Holmgren 2005b). The variation almost always does not correlate with geography, and the recognition of varieties in this species is neither practical nor very useful. All of those authors admitted that these varieties are “very weak at best” (Rollins, p. 554). Of them, perhaps var. pubicarpum (including var. elongatum) might merit recognition. It is distributed in almost all of the Mountain and Pacific states and is distinguished from the other varieties solely by the presence of trichomes or minute papillae on the fruit valves. The density of these trichomes ranges from moderate and covering the entire valve surface to very sparse and represented by individual papillate trichomes restricted to the valve margin. Furthermore, the length of these trichomes may vary from ca. 0.01 to 0.3 mm. In some species (e.g., L. dictyotum) both glabrous- and pubescent-fruited forms occur, yet none of the above authors gave formal recognition to both forms. It is not known if both glabrous and puberulent fruits occur within the same population in L. densiflorum. The species is autogamous, but nothing is known about the rates of gene flow between and within populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 579. FNA vol. 7, p. 577.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Synonyms L. densiflorum var. elongatum, L. densiflorum var. macrocarpum, L. densiflorum var. pubicarpum, L. densiflorum var. ramosum, L. elongatum, L. neglectum, L. pubicarpum, L. ramosum Thlaspi campestre, Neolepia campestris
Name authority Schrader: Index Seminum (Göttingen) 1832: 4. (1832) (Linnaeus) W. T. Aiton: in W. Aiton and W. T. Aiton, Hortus Kew. 4: 88. (1812)
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