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globe-pod hoary-cress, hairy whitetop, white-top

baby-seed pepper-weed, common pepper-grass, common pepper-weed, elongate peppergrass, hairy-fruit peppergrass, large-fruit peppergrass, miner's pepper-weed, prairie pepper-grass, prairie pepperweed

Habit Perennials; (rhizomatous); often densely hirsute. Annuals or biennials; puberulent or glabrous, (trichomes cylindrical).
Stems

simple or several from base, erect or ascending, branched distally, (1–)1.5–3.5(–5) dm.

simple from base, erect, branched distally, (1–)2.5–5(–6.5) dm.

Basal leaves

(often withered by anthesis);

not rosulate;

petiole 0.5–1.5 cm;

blade obovate to oblanceolate, (1–)2–6(–7) cm × 3–20 mm, margins dentate to sinuate.

(early withered); rosulate;

petiole 0.5–1.5(–2) cm;

blade oblanceolate, spatulate, or oblong, (1.5–)2.5–8(–11) cm × 5–10(–20) mm, margins coarsely serrate or pinnatifid.

Cauline leaves

sessile;

blade oblong or lanceolate, 1–5(–8) cm × (3–)5–15(–30) mm, base sagittate, margins dentate or subentire, (surfaces pubescent).

shortly petiolate;

blade narrowly oblanceolate or linear, (0.7–)1.3–6.2(–8) cm × (0.5–)1.5–10(–18) mm, base attenuate to cuneate, not auriculate, margins usually entire or irregularly serrate to dentate, rarely pinnatifid.

Racemes

(usually corymbose, rarely paniculate), rarely elongated in fruit;

rachis pubescent, trichomes often curved.

considerably elongated in fruit;

rachis puberulent, trichomes straight, slender to subclavate.

Flowers

sepals oblong, 1.4–2 × 0.7–1 mm;

petals white, broadly obovate, (2.2–)2.8–4 × 1–3 mm, claw 1–1.4 mm;

stamens 6;

filaments 2–2.5 mm, (glabrous);

anthers 0.4–0.5 mm.

sepals oblong, 0.5–0.8(–1) × 0.3–0.5 mm;

petals (absent or rudimentary) white, filiform, 0.3–0.9 mm, claw absent;

stamens 2, median;

filaments 0.6–1(–1.8) mm;

anthers 0.1–0.2 mm.

Fruiting pedicels

divaricate to ascending, straight or slightly curved, (terete), 3–9(–12) × 0.2–0.3 mm, pubescent.

divaricate-ascending to horizontal, straight or slightly recurved, (terete), (1.5–)2–3.5(–4) × 0.15–0.25 mm, puberulent adaxially.

Fruits

(indehiscent), globose or, rarely, subglobose, (2–)3–4.4(–5) mm diam., (inflated), apically not winged, apical notch absent;

valves thin, smooth, not veined, densely puberulent;

style 0.5–1.5 mm.

obovate to obovate-suborbicular, (2–)2.5–3(–3.5) × 1.5–2.5(–3) mm (widest beyond middle), apically winged, apical notch 0.2–0.4 mm deep;

valves thin, smooth, not veined, glabrous or sparsely puberulent (at least on margin);

style 0.1–0.2 mm, included in apical notch.

Seeds

(brown or dark brown), ovoid, 1.5–2 × 1–1.5 mm.

ovate, 1.1–1.3(–1.5) × 0.8–0.9 mm.

2n

= 16.

= 32.

Lepidium appelianum

Lepidium densiflorum

Phenology Flowering May–Sep. Flowering May–Jul.
Habitat Roadsides, sagebrush communities, alkaline meadows, waste grounds, ditch and stream sides, fields, pastures Waste places, disturbed sites, prairies, fields, pastures, grasslands, chaparral, meadows, sagebrush flats, flood plains, gravelly hillsides, rock crevices, seashores, sandy places, shaley barrens, open mesa, roadsides
Elevation 400-2400 m (1300-7900 ft) 0-3500 m (0-11500 ft)
Distribution
from FNA
CA; CO; ID; MI; MO; MT; ND; NE; NM; NV; OR; PA; UT; WA; WI; WY; AB; BC; MB; SK; c Asia [Introduced in North America; introduced also in South America, other parts of Asia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Mexico (Chihuahua, Nuevo León) [Introduced in Europe, Asia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Lepidium appelianum has become a noxious weed in most of its range in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

North American records of Lepidium apetalum Willdenow mostly represent misidentifications of L. densiflorum. The latter has obovate fruits widest beyond the middle, whereas L. apetalum has elliptic fruits widest at the middle.

The number and limits of the varieties recognized in Lepidium densiflorum, as well as the characters used to delimit them, vary among authors (A. Thellung 1906; C. L. Hitchcock 1936; G. A. Mulligan 1961; R. C. Rollins 1993; N. H. Holmgren 2005b). The variation almost always does not correlate with geography, and the recognition of varieties in this species is neither practical nor very useful. All of those authors admitted that these varieties are “very weak at best” (Rollins, p. 554). Of them, perhaps var. pubicarpum (including var. elongatum) might merit recognition. It is distributed in almost all of the Mountain and Pacific states and is distinguished from the other varieties solely by the presence of trichomes or minute papillae on the fruit valves. The density of these trichomes ranges from moderate and covering the entire valve surface to very sparse and represented by individual papillate trichomes restricted to the valve margin. Furthermore, the length of these trichomes may vary from ca. 0.01 to 0.3 mm. In some species (e.g., L. dictyotum) both glabrous- and pubescent-fruited forms occur, yet none of the above authors gave formal recognition to both forms. It is not known if both glabrous and puberulent fruits occur within the same population in L. densiflorum. The species is autogamous, but nothing is known about the rates of gene flow between and within populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 576. FNA vol. 7, p. 579.
Parent taxa Brassicaceae > tribe Lepidieae > Lepidium Brassicaceae > tribe Lepidieae > Lepidium
Sibling taxa
L. acutidens, L. alyssoides, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. densiflorum, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
L. acutidens, L. alyssoides, L. appelianum, L. austrinum, L. barnebyanum, L. campestre, L. chalepense, L. coronopus, L. crenatum, L. davisii, L. dictyotum, L. didymum, L. draba, L. eastwoodiae, L. flavum, L. fremontii, L. heterophyllum, L. huberi, L. integrifolium, L. jaredii, L. lasiocarpum, L. latifolium, L. latipes, L. montanum, L. nanum, L. nitidum, L. oblongum, L. ostleri, L. oxycarpum, L. papilliferum, L. paysonii, L. perfoliatum, L. pinnatifidum, L. ramosissimum, L. ruderale, L. sativum, L. sordidum, L. strictum, L. thurberi, L. tiehmii, L. virginicum
Synonyms Hymenophysa pubescens, Cardaria pubescens, Cardaria pubescens var. elongata L. densiflorum var. elongatum, L. densiflorum var. macrocarpum, L. densiflorum var. pubicarpum, L. densiflorum var. ramosum, L. elongatum, L. neglectum, L. pubicarpum, L. ramosum
Name authority Al-Shehbaz: Novon 12: 7. (2002) Schrader: Index Seminum (Göttingen) 1832: 4. (1832)
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