Lepidium alyssoides |
Lepidium tiehmii |
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mesa pepperwort |
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Habit | Perennials or subshrubs; (woody base often aboveground); glabrous or minutely puberulent. | Perennials; (caudex woody, to 1 cm diam., covered with persistent petiolar remains); glabrous throughout. |
Stems | few to several from base, erect to ascending, branched throughout, (0.7–)1–4.8(–6.1) dm. |
simple from base (caudex branch), erect, branched distally, 1–7.5 dm. |
Basal leaves | often not rosulate; petiole 1–6 cm; blade pinnately lobed, (1–)1.5–8(–11) cm × (5–)10–35 mm, margins (of lobes) entire or denticulate. |
rosulate; petiole (1.5–)2.5–10(–13) cm; blade oblong to lanceolate, (2.5–)4.5–9(–14) cm × 15–40 mm, margins entire. |
Cauline leaves | sessile; blade linear, (0.8–)1.3–7(–9.5) cm × (0.7–)1–2(–3) mm, base attenuate, not auriculate, margins entire. |
shortly petiolate; blade oblanceolate, (much smaller than basal), base attenuate-cuneate, not auriculate, margins entire. |
Racemes | elongated in fruit; rachis puberulent or glabrous. |
(paniculate), considerably elongated in fruit. |
Flowers | sepals ovate to oblong, 1–2 × 0.8–1 mm; petals white, suborbicular, 2–3 × 1–2 mm, claw 0.5–1.5 mm; stamens 6; filaments 1.5–2 mm, (glabrous); anthers 0.2–0.4 mm. |
sepals oblong, 2.5–3(–4) × 1.2–1.8 mm; petals creamy white to pale yellow, obovate to oblanceolate, 4–5.5(–6.5) × 2–3.3 mm, claw 1.5–2.2 mm; stamens 6; filaments (median pairs) 3.5–4.5 mm; anthers 1–1.2 mm. |
Fruiting pedicels | divaricate to horizontal, straight or recurved to somewhat sigmoid, (terete), 3.5–8(–11) × 0.2 mm, glabrous or puberulent adaxially. |
divaricate-ascending, straight, (terete), 8–15 × 0.4–0.5 mm. |
Fruits | broadly ovate, 2–3.7(–4.3) × (1.5–)1.8–2.9(–3.4) mm, apically winged, apical notch 0.1–0.3(–0.4) mm deep; valves thin, smooth, not veined, glabrous; style 0.2–0.6 mm, exserted beyond apical notch. |
obovate to somewhat rhomboid, 7–11 × 5–6.5 mm, apically not winged, apical notch absent; valves thin, smooth, obscurely veined; style 0.2–0.6 mm. |
Seeds | ovate, 1.5–1.8(–2) × 0.9–1.2(–1.5) mm. |
oblong, 3.7–4.5 × 1.6–2 mm. |
2n | = 32. |
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Lepidium alyssoides |
Lepidium tiehmii |
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Phenology | Flowering May–Jul. | Flowering Apr–Jun. |
Habitat | Pinyon-juniper or sagebrush communities, prairies, grasslands, sandstone outcrops, gypsum flats, sand dunes, dry flats and river bottoms, gravelly roadsides | Rocky crevices and slopes in sagebrush communities |
Elevation | 1200-2800 m (3900-9200 ft) | 1400-1800 m (4600-5900 ft) |
Distribution |
AZ; CO; NM; NV; TX; UT; WY; Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí)
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NV |
Discussion | Of the five varieties of Lepidium alyssoides recognized by R. C. Rollins (1993), one (var. mexicanum Rollins) is a short-tufted form of the species restricted to Mexico that does not seem to merit recognition, another (var. junceum) is a glabrescent form of the type variety, a third (var. eastwoodiae) is treated below as a distinct species, and the fourth (var. angustifolium) is included here within L. alyssoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Lepidium tiehmii is known from mountain ranges in Douglas and Lyon counties. It was described and has been maintained (N. H. Holmgren 2005b) in Stroganowia, a genus now united with Lepidium (I. A. Al-Shehbaz et al. 2002) that otherwise is disjunct and restricted to the central Asian states of the Former Soviet Union and adjacent western China. In our opinion, the similarity of this species to those Asian ones formerly placed in Stroganowia is superficial and is the result of convergence rather than descent. The cotyledonary type was erroneously reported as conduplicate (R. C. Rollins 1993; N. H. Holmgren 2005b). In the several seeds that we dissected it was always incumbent. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 575. | FNA vol. 7, p. 594. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | L. alyssoides var. angustifolium, L. alyssoides var. junceum, L. alyssoides var. minus, L. alyssoides var. polycarpum, L. alyssoides var. streptocarpum, L. montanum subsp. alyssoides, L. montanum var. alyssoides, L. montanum subsp. angustifolium, L. montanum var. angustifolium, L. tortum | Stroganowia tiehmii |
Name authority | A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 10. (1849) | (Rollins) Al-Shehbaz: Novon 12: 9. (2002) |
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