Lepidium alyssoides |
Lepidium didymum |
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mesa pepperwort |
lesser pepper-grass, lesser swine-cress, lesser wart-cress |
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Habit | Perennials or subshrubs; (woody base often aboveground); glabrous or minutely puberulent. | Annuals; (fetid); glabrous or pilose. |
Stems | few to several from base, erect to ascending, branched throughout, (0.7–)1–4.8(–6.1) dm. |
few to several from base, erect to ascending or decumbent, branched distally, 1–4.5(–7) dm. |
Basal leaves | often not rosulate; petiole 1–6 cm; blade pinnately lobed, (1–)1.5–8(–11) cm × (5–)10–35 mm, margins (of lobes) entire or denticulate. |
(soon withered); not rosulate; petiole 0.5–4(–6) cm; blade 1- or 2-pinnatisect, 1–6(–8) cm, margins (of lobes) entire or dentate (sometimes deeply lobed). |
Cauline leaves | sessile; blade linear, (0.8–)1.3–7(–9.5) cm × (0.7–)1–2(–3) mm, base attenuate, not auriculate, margins entire. |
shortly petiolate to subsessile; blade similar to basal, smaller and less divided distally, lobes lanceolate to oblong or elliptic, 1.5–3.5(–4.5) cm ×5–12 mm, base not auriculate, margins (of lobes) entire, serrate, or incised. |
Racemes | elongated in fruit; rachis puberulent or glabrous. |
elongated in fruit; rachis glabrous or pubescent, trichomes straight, cylindrical. |
Flowers | sepals ovate to oblong, 1–2 × 0.8–1 mm; petals white, suborbicular, 2–3 × 1–2 mm, claw 0.5–1.5 mm; stamens 6; filaments 1.5–2 mm, (glabrous); anthers 0.2–0.4 mm. |
sepals (tardily deciduous), ovate, 0.5–0.7(–0.9) mm; petals white, elliptic to linear, 0.4–0.5 × ca. 0.1 mm, claw absent; stamens 2, median; filaments 0.3–0.6 mm; anthers 0.1–0.2 mm. |
Fruiting pedicels | divaricate to horizontal, straight or recurved to somewhat sigmoid, (terete), 3.5–8(–11) × 0.2 mm, glabrous or puberulent adaxially. |
divaricate to horizontal, straight slightly recurved, (terete), 1.4–2.5(–4) × 0.15–2 mm, glabrous or sparsely pubescent adaxially. |
Fruits | broadly ovate, 2–3.7(–4.3) × (1.5–)1.8–2.9(–3.4) mm, apically winged, apical notch 0.1–0.3(–0.4) mm deep; valves thin, smooth, not veined, glabrous; style 0.2–0.6 mm, exserted beyond apical notch. |
schizocarpic, didymous, 1.3–1.7 × 2–2.5 mm, apically not winged, apical notch 0.2–0.4 mm deep; valves thick, rugose, strongly veined, glabrous; style absent or obsolete, included in apical notch. |
Seeds | ovate, 1.5–1.8(–2) × 0.9–1.2(–1.5) mm. |
ovate, 1–1.2 × 0.7–0.8 mm. |
2n | = 32. |
= 32. |
Lepidium alyssoides |
Lepidium didymum |
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Phenology | Flowering May–Jul. | Flowering Mar–Jul. |
Habitat | Pinyon-juniper or sagebrush communities, prairies, grasslands, sandstone outcrops, gypsum flats, sand dunes, dry flats and river bottoms, gravelly roadsides | Roadsides, waste areas, lawns, pastures, fields, gardens, disturbed areas |
Elevation | 1200-2800 m (3900-9200 ft) | 0-1000 m (0-3300 ft) |
Distribution |
AZ; CO; NM; NV; TX; UT; WY; Mexico (Chihuahua, Coahuila, Nuevo León, San Luis Potosí)
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AL; AR; AZ; CA; CT; FL; GA; LA; MA; MD; ME; MO; MS; NC; NJ; NY; OH; OR; PA; SC; TN; TX; VA; WI; WV; BC; NB; NF; NS; QC; South America [Introduced in North America; introduced also in Mexico (Sinaloa), Central America (Honduras), Europe, Asia, s Africa, Australia]
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Discussion | Of the five varieties of Lepidium alyssoides recognized by R. C. Rollins (1993), one (var. mexicanum Rollins) is a short-tufted form of the species restricted to Mexico that does not seem to merit recognition, another (var. junceum) is a glabrescent form of the type variety, a third (var. eastwoodiae) is treated below as a distinct species, and the fourth (var. angustifolium) is included here within L. alyssoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 575. | FNA vol. 7, p. 580. |
Parent taxa | Brassicaceae > tribe Lepidieae > Lepidium | Brassicaceae > tribe Lepidieae > Lepidium |
Sibling taxa | ||
Synonyms | L. alyssoides var. angustifolium, L. alyssoides var. junceum, L. alyssoides var. minus, L. alyssoides var. polycarpum, L. alyssoides var. streptocarpum, L. montanum subsp. alyssoides, L. montanum var. alyssoides, L. montanum subsp. angustifolium, L. montanum var. angustifolium, L. tortum | Carara didyma, Coronopus didymus, Senebiera didyma, Senebiera incisa, Senebiera pinnatifida |
Name authority | A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 10. (1849) | Linnaeus: Syst. Nat. ed. 12, 2: 433. (1767): Mant Pl. 1: 92. (1767) |
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