Layia pentachaeta |
Layia glandulosa |
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Sierra layia, Sierra tidy tips |
white layia, white-daisy tidytips |
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Habit | Plants 5–100 cm (self-incompatible); glandular, strongly lemon- or acrid-scented. | Plants 3–60 cm (self-incompat-ible); glandular, often not strongly scented, sometimes spice-scented. | ||||
Stems | not purple-streaked. |
not purple-streaked (often uniformly dark purple). |
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Leaf | blades oblanceolate or lanceolate to linear, 6–110 mm, margins (basal leaves) 1–2-pinnatifid. |
blades obovate to linear, 6–100 mm, margins (basal leaves) toothed to lobed. |
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Involucres | ± hemispheric, 5–12 × 2–12+ mm. |
± campanulate to hemispheric, 4–11 × 3–11+ mm. |
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Ray florets | 3–14; laminae yellow or white, 3–26 mm. |
3–14; laminae often white, sometimes yellow or pale yellow, 3–22 mm. |
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Disc florets | 7–120+; corollas 3–6 mm; anthers yellow to brownish. |
17–100+; corollas 3.5–6.5 mm; anthers yellow to brownish. |
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Phyllaries | 4–14, apices shorter or longer than folded bases. |
3–14, apices often shorter (sometimes longer) than folded bases. |
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Ray cypselae | glabrous. |
glabrous. |
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Paleae | in 1 series between ray and disc florets. |
in 1 series between ray and disc florets. |
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Disc | pappi 0 or of 1–22 whitish, ± equal, setiform scales 1.5–3.5 mm, each proximally plumose, seldom adaxially woolly. |
pappi of 10–15 usually white (rarely tawny), linear-attenuate to subulate, ± equal scales 2–5 mm, each proximally plumose and often adaxially woolly. |
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2n | = 16. |
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Layia pentachaeta |
Layia glandulosa |
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Phenology | Flowering Feb–Jul. | |||||
Habitat | Openings in scrub, woodlands, forests, grasslands, and meadows, gravelly or sandy soils, sometimes dunes | |||||
Elevation | 0–2700 m (0–8900 ft) | |||||
Distribution |
CA
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AZ; CA; ID; NM; OR; UT; WA; Mexico (Baja California)
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Discussion | Subspecies 2 (2 in the flora). Molecular phylogenetic data have indicated that Layia pentachaeta is closely related to L. glandulosa and L. discoidea (B. G. Baldwin, unpubl.). Distributions of L. pentachaeta and L. glandulosa overlap broadly; the two species occur in different habitats and natural hybridization has been rarely detected (first-generation hybrids are of low to moderate fertility; J. Clausen 1951). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Layia glandulosa occurs in deserts of western North America, extending to the Pacific coast in central and southern California. As treated here (provisionally) and previously, L. glandulosa corresponds to a paraphyletic group; molecular phylogenetic data have indicated that L. discoidea is most closely related to a subset of lineages in L. glandulosa, including yellow-rayed populations previously recognized as subsp. lutea or var. lutea (B. G. Baldwin, unpubl.). Report of L. glandulosa from British Columbia has not been confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 266. | FNA vol. 21, p. 267. | ||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Madiinae > Layia | Asteraceae > tribe Heliantheae > subtribe Madiinae > Layia | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Blepharipappus glandulosa, L. glandulosa subsp. lutea, L. glandulosa var. lutea | |||||
Name authority | A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 108. (1857) | (Hooker) Hooker & Arnott: Bot. Beechey Voy., 358. (1839) | ||||
Web links |