Layia pentachaeta |
Asteraceae tribe Heliantheae subtribe Madiinae |
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Sierra layia, Sierra tidy tips |
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Habit | Plants 5–100 cm (self-incompatible); glandular, strongly lemon- or acrid-scented. | Annuals, perennials, subshrubs, or shrubs, (0.5–)1–250 cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | not purple-streaked. |
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Leaves | blades oblanceolate or lanceolate to linear, 6–110 mm, margins (basal leaves) 1–2-pinnatifid. |
mostly basal, or basal and cauline, or mostly cauline; opposite and/or alternate; usually sessile, rarely petiolate; blades usually spatulate to oblanceolate or linear to linear-elliptic, sometimes pinnately lobed, ultimate margins entire or toothed, faces usually hairy and glandular (glands sometimes elaborate, apices sometimes gland- or spine-tipped). |
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Involucres | ± hemispheric, 5–12 × 2–12+ mm. |
campanulate, cylindric, ellipsoid, globose, fusiform, hemispheric, obconic, or urceolate. |
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Receptacles | flat or convex to conic, ± paleate (paleae either in 1 series interior to ray florets, then usually connate and persistent, or 1 palea subtending each disc floret, then usually distinct and soon falling, or, in discoid heads, paleae functioning as phyllaries in 1 series and together constituting an “involucre”). |
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Ray florets | 3–14; laminae yellow or white, 3–26 mm. |
0, or 1–75+, pistillate, fertile; corollas usually yellow or whitish, sometimes proximally yellow and distally whitish, often marked with red or purple or turning red on drying, rarely orange or red-orange (apices usually ± deeply divided, lobes usually 3, sometimes 2). |
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Disc florets | 7–120+; corollas 3–6 mm; anthers yellow to brownish. |
1–200+, bisexual and fertile, or functionally staminate; corollas usually yellow, sometimes brown, orange, reddish, or whitish, tubes much shorter than to about equaling cylindric to funnelform throats, lobes 5, deltate; anther thecae usually dark (reddish to purple, often described as “black”), sometimes pale (yellow to brown); stigmatic papillae in 2 lines. |
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Phyllaries | 4–14, apices shorter or longer than folded bases. |
persistent or falling, 1–75+ in 1(–2) series or 0 (the “involucre” then interpreted as constituted of 1 series of receptacular paleae; phyllaries distinct or weakly connate, narrowly ovate to lanceolate or linear, equal or subequal, herbaceous, usually conduplicate, often each partly or wholly investing a subtended floret, apices often attenuate, faces usually glandular). |
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Calyculi | usually 0, sometimes 2–12+ bractlets (Centromadia, some members of Hemizonia and Lagophylla). |
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Heads | usually radiate, rarely discoid, usually in corymbiform, glomerulate, paniculiform, spiciform, or ± umbelliform arrays, sometimes borne singly. |
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Cypselae | obcompressed, compressed, or terete, often obpyramidal, clavate, or fusiform, glabrous or hairy (ray and disc often different); pappi 0, or persistent or falling, of awns, bristles, or scales (sometimes in combination) in 1–2 series (the elements often ciliate or plumose). |
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Ray cypselae | glabrous. |
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Paleae | in 1 series between ray and disc florets. |
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Disc | pappi 0 or of 1–22 whitish, ± equal, setiform scales 1.5–3.5 mm, each proximally plumose, seldom adaxially woolly. |
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Layia pentachaeta |
Asteraceae tribe Heliantheae subtribe Madiinae |
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Distribution |
CA
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HI; South America; Mostly w North America |
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Discussion | Subspecies 2 (2 in the flora). Molecular phylogenetic data have indicated that Layia pentachaeta is closely related to L. glandulosa and L. discoidea (B. G. Baldwin, unpubl.). Distributions of L. pentachaeta and L. glandulosa overlap broadly; the two species occur in different habitats and natural hybridization has been rarely detected (first-generation hybrids are of low to moderate fertility; J. Clausen 1951). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 24, species 121 (20 genera, 83 species in the flora). Madiinae has long included the “tarweeds,” which are centered in or restricted to the California Floristic Province, the silversword alliance of the Hawaiian Islands (Argyroxiphium, Dubautia, and Wilkesia), and Raillardella (in a broad sense), which was formerly included in Senecioneae (Carlquist et al. 2003). Such a circumscription was used by H. Robinson (1981) and is followed here. In papers by B. G. Baldwin and B. Wessa (2000) and Baldwin et al. (2002), Madiinae were linked to genera traditionally included in Heliantheae (e.g., Eriophyllum, Hulsea, Lasthenia, Venegasia) and in Senecioneae (e.g., Arnica) and Madiinae plus Arnicinae, Baeriinae, Hulseinae, and Venegasiinae were included in tribe Madieae. Key to genera of Madiinae here is based on key by B. G. Baldwin (in S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 266. | FNA vol. 21, p. 254. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Madiinae > Layia | Asteraceae > tribe Heliantheae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 108. (1857) | Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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