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hawkweed layia, tall tidytips

Sierra layia, Sierra tidy tips

Habit Plants 5–130 cm (self-compatible); glandular, usually strongly sweet- or pungent-scented. Plants 5–100 cm (self-incompatible); glandular, strongly lemon- or acrid-scented.
Stems

purple-streaked.

not purple-streaked.

Leaf

blades elliptic, lanceolate, linear, or oblanceolate, 7–150 mm, margins (basal leaves) lobed to toothed.

blades oblanceolate or lanceolate to linear, 6–110 mm, margins (basal leaves) 1–2-pinnatifid.

Involucres

± ellipsoid to obconic, 4–9 × 4–9+ mm.

± hemispheric, 5–12 × 2–12+ mm.

Ray florets

6–16;

laminae yellow, 1–4 mm.

3–14;

laminae yellow or white, 3–26 mm.

Disc florets

9–80;

corollas 2.5–4.5 mm;

anthers ± dark purple.

7–120+;

corollas 3–6 mm;

anthers yellow to brownish.

Phyllaries

6–16, apices shorter than folded bases.

4–14, apices shorter or longer than folded bases.

Ray cypselae

glabrous.

glabrous.

Paleae

in 1 series between ray and disc florets.

in 1 series between ray and disc florets.

Disc

pappi of 10–16 white to rufous or purplish, ± equal bristles or setiform scales 2–4 mm, each proximally plumose, not adaxially woolly.

pappi 0 or of 1–22 whitish, ± equal, setiform scales 1.5–3.5 mm, each proximally plumose, seldom adaxially woolly.

2n

= 16, 32.

Layia hieracioides

Layia pentachaeta

Phenology Flowering Apr–Jul.
Habitat Open, often grassy, or semishady sites in chaparral, woodlands, forests, and coastal scrub, often sandy soils, disturbed sites
Elevation 0–1200 m (0–3900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Layia hieracioides occurs in the southern San Francisco Bay area, South Coast Ranges, and Western Transverse Ranges. Diploids and tetraploids are morphologically similar and reportedly geographically distinct, with diploids documented from the San Francisco Bay area and tetraploids (= L. paniculata) documented to the south. Natural, ± sterile hybrids between tetraploid plants and L. glandulosa have been documented from the Central Coast (B. D. Tanowitz and J. W. Adams 1986; R. F. Hoover 8369, UC).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Molecular phylogenetic data have indicated that Layia pentachaeta is closely related to L. glandulosa and L. discoidea (B. G. Baldwin, unpubl.). Distributions of L. pentachaeta and L. glandulosa overlap broadly; the two species occur in different habitats and natural hybridization has been rarely detected (first-generation hybrids are of low to moderate fertility; J. Clausen 1951).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ray corollas yellow
subsp. pentachaeta
1. Ray corollas white
subsp. albida
Source FNA vol. 21, p. 268. FNA vol. 21, p. 266.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Madiinae > Layia Asteraceae > tribe Heliantheae > subtribe Madiinae > Layia
Sibling taxa
L. carnosa, L. chrysanthemoides, L. discoidea, L. fremontii, L. gaillardioides, L. glandulosa, L. heterotricha, L. jonesii, L. leucopappa, L. munzii, L. pentachaeta, L. platyglossa, L. septentrionalis
L. carnosa, L. chrysanthemoides, L. discoidea, L. fremontii, L. gaillardioides, L. glandulosa, L. heterotricha, L. hieracioides, L. jonesii, L. leucopappa, L. munzii, L. platyglossa, L. septentrionalis
Subordinate taxa
L. pentachaeta subsp. albida, L. pentachaeta subsp. pentachaeta
Synonyms Madaroglossa hieracioides, L. paniculata
Name authority (de Candolle) Hooker & Arnott: Bot. Beechey Voy., 358. (1839) A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 108. (1857)
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