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hairy goldfields, maritime goldfields, maritime or seaside goldfields, seaside goldfields

Habit Annuals, to 25 cm (herbage not sweetly scented). Annuals, perennials, subshrubs, or shrubs, 1–200 cm.
Stems

usually prostrate or decumbent, rarely erect, branched proximally, glabrous or hairy at nodes and distally.

Leaves

linear to oblanceolate, 10–90 × 2–12 mm, (fleshy) margins entire or lobed, faces glabrous.

basal, basal and cauline, or mostly cauline; mostly opposite (Lasthenia) or mostly alternate; usually sessile, sometimes obscurely petiolate;

blades (often 1–2 times pinnately lobed) or lobes often linear, ultimate margins entire or toothed, faces often ± woolly to tomentose, sometimes glabrate or glabrous, often gland-dotted.

Involucres

hemispheric, 4–7 mm.

ovoid or obconic to campanulate or hemispheric.

Receptacles

conic, muricate, glabrous.

flat, convex, hemispheric, or conic (smooth, knobby, or pitted, glabrous or hairy), usually epaleate (paleae usually 0, rare in Eriophyllum).

Ray florets

7–12; (corollas light to golden yellow) laminae oblong, 1–3 mm.

0 or 4–21, pistillate, fertile (3–8 peripheral florets pistillate, fertile, corollas tubular in Amblyopappus and Monolopia congdonii);

corollas yellow to orange, often darker proximally, sometimes purplish (usually ± bilabiate in Monolopia).

Disc florets

2–300, bisexual, fertile;

corollas yellow to orange, tubes shorter than or about equaling funnelform or campanulate throats, lobes 4–5, deltate, glabrous or papillate;

anther thecae usually pale;

stigmatic papillae in 2 lines.

Phyllaries

6–14, lanceolate to ovate, hairy (especially at margins and midribs).

persistent, mostly 3–18 in 1–2 series, (erect or reflexed in fruit) distinct or connate, mostly elliptic, lanceolate, ovate, or obovate, usually ± equal, mostly herbaceous, sometimes indurate (at least proximally), flat or weakly cupped at bases, sometimes scarious-margined, often woolly to tomentose, sometimes glabrate or glabrous.

Calyculi

0.

Heads

radiate, discoid, or disciform, borne singly or in corymbiform, glomerate, or paniculiform arrays.

Cypselae

gray, linear to narrowly clavate, (2–)2.5–3 mm, ± hairy;

pappi usually of 4–6(–12) brown, lanceolate or subulate, aristate scales plus 4–5+ shorter, laciniate scales, rarely 0.

clavate or obovoid to terete, or obpyramidal, sometimes compressed or obcompressed, glabrous, hairy, or papillate (compressed, callous-margined, and ciliolate in Eatonella, Lasthenia chrysantha, and Monolopia congdonii; sometimes winged in Monolopia);

pappi 0 or of 1–12+ aristate, erose, laciniate, or truncate scales or awns in 1–2 series (often 2 sorts of scales in combination on 1 cypsela).

Anther

appendages ± oblong, obtuse (style apices ± deltate, glabrous or with apical tufts of hairs and subapical fringes of shorter hairs).

2n

= 8.

Lasthenia maritima

Asteraceae tribe Heliantheae subtribe Baeriinae

Phenology Flowering May–Jul.
Habitat Seabird roosting sites, coastal headlands, offshore rocks, islands
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
w North America; Mexico; w South America
Discussion

Lasthenia maritima is a self-pollinating, “guano endemic” of seabird nesting grounds. It is typically found on offshore islands and rocks from the Farallon Islands, California, to the northern tip of Vancouver Island, British Columbia and rarely occurs on the mainland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 9, species 44 (7 genera, 41 species in the flora).

Members of Baeriinae are found mostly in western North America; there are disjuncts in western South America. H. Robinson (1981) treated Baeriinae as a relatively isolated element among epaleate subtribes of Heliantheae. B. G. Baldwin (in Baldwin et al. 2002) included Baeriinae within Madieae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves all or mostly opposite
Lasthenia
1. Leaves mostly alternate (proximal sometimes opposite)
→ 2
2. Leaves glabrous (often granular-glandular, not woolly)
Amblyopappus
2. Leaves ± woolly or tomentose (usually stems and/or phyllaries as well)
→ 3
3. Heads ± disciform (4–7 peripheral florets pistillate, corollas with laminae ca. 0.5 mm)
Monolopia
3. Heads usually radiate, sometimes discoid
→ 4
4. Phyllaries usually reflexed in fruit (ray laminae 2–2.5 mm; pappi of 2, ± lacini-ate to nearly entire, often uniaristate scales 1.5–3 mm)
Eatonella
4. Phyllaries usually ± erect in fruit
→ 5
5. Ray florets 7–11 (corollas obscurely bilabiate, laminae each opposed by adaxial lobe or tooth)
Monolopia
5. Ray florets 0 or 3–13 (corollas not bilabiate)
→ 6
6. Annuals; ray cypselae obcompressed (disc corollas with rings of hairs at bases of limbs)
Pseudobahia
6. Annuals, perennials, subshrubs, or shrubs; ray cypselae usually prismatic, 4–5-angled (disc corollas without rings of hairs)
→ 7
7. Annuals, perennials, subshrubs, or shrubs; phyllaries 4–13(–15) in 1+ series; pappi 0, or coroniform, or of 6–12+ (distinct) scales (in 1–2 similar or contrasting series)
Eriophyllum
7. Subshrubs; phyllaries 8–16+ in ± 2 series; pappi of 2–6+ (unequal or 2 opposite, longer, and ± equal, basally connate, oblong to subulate) scales (tips acute to erose)
Constancea
Source FNA vol. 21, p. 347. FNA vol. 21, p. 335.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Baeriinae > Lasthenia > sect. Ptilomeris Asteraceae > tribe Heliantheae
Sibling taxa
L. burkei, L. californica, L. chrysantha, L. conjugens, L. coronaria, L. debilis, L. ferrisiae, L. fremontii, L. glaberrima, L. glabrata, L. gracilis, L. leptalea, L. microglossa, L. minor, L. ornduffii, L. platycarpha
Subordinate taxa
Amblyopappus, Constancea, Eatonella, Eriophyllum, Lasthenia, Monolopia, Pseudobahia
Synonyms Burrielia maritima, Baeria maritima, Baeria minor subsp. maritima, L. minor subsp. maritima subtribe Eriophyllinae
Name authority (A. Gray) M. C. Vasey: Madroño 32: 139. (1985) Bentham & Hooker f.: Gen. Pl. 2: 200. (1873)
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