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scurf-pea

dune scurfpea, lance-leaf scurf-pea, lemon scurfpea, wild lemonweed

Habit Herbs, perennial, or subshrubs, unarmed; roots deep and woody or rhizomatous. Herbs.
Stems

erect or spreading, green, smooth or lightly striate, sometimes distinctly ribbed apically, branched, basally often with thin, tan to brown cataphylls, glandular, glabrate, or sericeous to densely pubescent throughout or basally, sometimes glabrate.

branched throughout, not rushlike.

Leaves

alternate, usually palmately compound, rarely pseudopalmate, often deciduous by anthesis except basally;

stipules present, caducous or tardily deciduous, usually adnate to petiole, rarely connate behind petiole, often modified to form cataphylls at base of stem, linear-lanceolate;

petiolate, petiole not jointed to stem;

leaflets (1 or)3 or 5, persistent or caducous, sometimes absent at maturity, blade margins entire, surfaces strigose, glabrate, or sparsely pubescent.

usually palmate, rarely pseudopalmate, persistent through flowering;

petiole (0.5–)0.9–2.9 cm;

leaflets 3(or 5), blades obovate or oblanceolate to linear, 17–35(–40) × 2–13(–16) mm, base attenuate, apex retuse to acuminate, surfaces glabrate to sparsely pubescent.

Inflorescences

1–3(or 4)-flowered per node, indeterminate pseudoracemes, with 3–22 nodes;

bracts caducous, lanceolate to rhombic.

rachis 0.5–14 cm, internodes to 1.5 cm, elongating or not in fruit.

Peduncles

0.7–12 cm.

Flowers

papilionaceous;

calyx campanulate, lobes 5, lobes pallid green or green throughout, triangular, usually less than 1/2 length of calyx tube, mostly equal in length, abaxial lobe sometimes slightly longer, flaring back and tearing along a lateral sinus in fruit, glandular, pubescent or glabrate to sericeous;

corolla violet to blue, purple, white, or bicolored;

stamens 10, diadelphous, distinct portion of filaments filiform;

anthers basifixed.

calyx 1.5–2.5 mm;

corolla blue to purple, white, or bicolored, 5–7 mm.

Fruits

legumes, deciduous above receptacle, sessile, subglobose, indehiscent, pilose, strigose, or glabrate.

Legumes

conspicuously glandular, sparsely strigose to glabrate.

Seed

1, brown, somewhat compressed, round to elliptic, smooth.

x

= 11.

2n

= 22.

Ladeania

Ladeania lanceolata

Phenology Flowering early spring–late summer.
Habitat Aeolian sand dunes, sandsage and sand prairies.
Elevation 20–1800 m. (100–5900 ft.)
Distribution
North America
[BONAP county map]
from FNA
AZ; CA; CO; IA; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; SK
[BONAP county map]
Discussion

Species 2 (2 in the flora).

Recent treatments of Psoralidium Rydberg have recognized three species: the narrow endemic, P. junceum, and two widespread species, P. tenuiflorum and P. lanceolatum (J. W. Grimes 1990; D. Isely 1998). A phylogenetic analysis of Psoralidium and other genera of tribe Psoraleeae based on eight DNA regions found the type of the genus, P. tenuiflorum, to be nested within Pediomelum Rydberg (A. N. Egan and K. A. Crandall 2008), resulting in the transfer of P. tenuiflorum to Pediomelum as P. tenuiflorum (Pursh) A. N. Egan. Because the type of Psoralidium was moved to Pediomelum, the new genus Ladeania was created to accommodate the remaining two species, with the type designated as L. lanceolata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ladeania lanceolata is variable in morphology, which has resulted in the recognition of three to five taxa (P. A. Rydberg 1919; S. L. Welsh et al. 2015). The length of inflorescences, vestiture of pods, and shape of leaflets vary along geographic trendlines throughout its range and seem to grade from one trend to another.

The names Psoralea lanceolata Pursh (1813) and P. elliptica Pursh (1813) have equal priority. However, J. Torrey and A. Gray (1838–1843) placed P. elliptica in synonymy under P. lanceolata, establishing priority of the latter name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Mature stems rushlike, leafless or with few basal leaves when flowering; peduncles 15–40+ cm; calyces 2.5–4 mm; legumes eglandular.
L. juncea
1. Mature stems not rushlike, leaves persistent through flowering; peduncles 0.7–12 cm; calyces 1.5–2.5 mm; legumes glandular.
L. lanceolata
Source FNA vol. 11. Author: Ashley N. Egan. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae Fabaceae > subfam. Faboideae > Ladeania
Sibling taxa
L. juncea
Subordinate taxa
L. juncea, L. lanceolata
Synonyms Psoralea lanceolata, Lotodes elliptica, P. elliptica, P. micrantha, P. scabra, P. stenophylla, P. stenostachys, Psoralidium lanceolatum
Name authority A. N. Egan & Reveal: Novon 19: 311, fig. 1. (2009) (Pursh) A. N. Egan & Reveal: Novon 19: 312. (2009)
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